Cortaderia boliviensis M.Lyle, Novon 6(1): 72. 1996.

Testoni, Daniel & Linder, H. Peter, 2017, Synoptic taxonomy of Cortaderia Stapf (Danthonioideae, Poaceae), PhytoKeys 76, pp. 39-69 : 57

9. Cortaderia boliviensis M.Lyle, Novon 6(1): 72. 1996. Fig. 3C

Cortaderia bifida Pilg. var. grandiflora Henrard, Meded. Rijks-Herb. 40: 67. 1921. Type: Bolivia, Departamento Cochabamba: "Charactergrass der Andenwiesen über Tablas, feuchte Stellen, 3400 m, Mai 1911, T. C. J. Herzog 2194 (holotype: L; isotypes: S, US!, W!, Z!).

Etymology.

- ense (Latin), denoting origin. From Bolivia.

Taxonomy.

This species is very similar to Cortaderia nitida , with which it shares the (usually) non-lacerated, entire leaf sheaths and the shape of the lemmas, as well as largely similar leaf anatomy. However, neither chloroplast nor nuclear genome indicates such a relationship for Cortaderia boliviensis ( Pirie et al. 2009). It differs by the horizontally shattering sheaths. More inconsistent differences are in the indumentum of the floret, with the callus indumentum of Cortaderia boliviensis being shorter than in Cortaderia nitida . Lyle (1996) diagnosed Cortaderia boliviensis against Cortaderia bifida , under which it was originally described as a variety by Henrard in 1921. Mostly it is very different from Cortaderia bifida : the latter has much longer lemma setae and the basal sheaths are lacerated and not shattered. The type collection, however, is easily confused with Cortaderia bifida due to the long awns and setae, and somewhat fragmented leaf sheaths. The leaf anatomy is also quite different.

The leaf anatomy (Fig. 3E ) is like that of Cortaderia nitida , with adaxial grooves and a well-developed abaxial collenchyma layer. There are differences in detail, and wider sampling may well indicate that this is within-species variation.

Figures

Figure 3. Leaf anatomy of Cortaderia, as evident from transverse sections. A Cortaderia egmontiana (Moore 2677) B Cortaderia modesta (Glaziou 17913) C Cortaderia hieronymi (Garcia 563) D Cortaderia nitida (Laegaard 53121) E Cortaderia boliviensis (Beck 11273); F Cortaderia sericantha (Ramsay 967); G Cortaderia echinata (Peterson 21587) H Cortaderia bifida (Renvoize 4202) I Cortaderia hapalotricha (Laegaard 53305) J Cortaderia roraimensis (Maguire 60448). Structures referred to in the descriptions are labelled as follows: 1, multi-layered abaxial sub-epidermal collenchyma layer; 2, adaxial islands of collenchyma in the abaxial grooves; 3, chlorenchyma; 4, primary vascular bundle; 5, midrib.