Muellerpalia haszprunari, Neubauer, Thomas A., Harzhauser, Mathias, Kroh, Andreas, Elisavet, Georgopoulou & Mandic, Oleg, 2014

Neubauer, Thomas A., Harzhauser, Mathias, Kroh, Andreas, Elisavet, Georgopoulou & Mandic, Oleg, 2014, Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda. Part 2, ZooKeys 429, pp. 13-46 : 18-20

Taxon classification Animalia ORDO FAMILIA

Muellerpalia haszprunari nom. n.

Valvata simplex Fuchs, 1870 in Fuchs 1870b: 535, pl. 21, figs 4-6 [non Valvata tricarinata var. simplex Gould, 1841].

Valvata (Valvata) simplex simplex Fuchs; Wenz 1928a: 2474 (cum syn.) [non Gould 1841].

Valvata simplex Fuchs; Strausz 1942: 80 [non Gould 1841].

Hauffenia simplex (Fuchs); Schlickum 1978: 247, pl. 18, fig. 3 [non Gould 1841].

Hauffenia simplex (Fuchs 1870); Harzhauser and Binder 2004: 9 [non Gould 1841].

Etymology.

In honor of Gerhard Haszprunar (Bavarian State Collection of Zoology Munich and Ludwig Maximilians University Munich), who summarized all existing names of living and fossil valvatids in a comprehensive nomenclator ( Haszprunar 2014).

Type locality.

Tihany at Lake Balaton, Veszprém, Hungary.

Age.

Late Miocene (Late Pannonian, Transdanubian sensu Sacchi and Horváth 2002; Sztanó et al. 2013).

Type material.

According to the inventory books of the Natural History Museum Vienna the material should be stored there, but despite great effort it could not be located.

Discussion.

This species is a primary homonym of the extant taxon Valvata tricarinata var. simplex Gould, 1841 (p. 226) from Massachusetts, USA. The American taxon was elevated to species level by Fluck (1932). As the European species was combined with various genera since its first description and several subspecies have been described, a summary of its history is given below.

Already Schlickum (1978) considered Valvata simplex Fuchs, 1870 to belong to the Hydrobiidae and placed it in the genus Hauffenia , based on similarities of morphology and size. Recently, Bandel (2010) introduced the new genus Muellerpalia for Valvata bicincta Fuchs, 1870 in Fuchs (1870b), Valvata carinata Fuchs, 1870 in Fuchs (1870b), Planorbis varians Fuchs, 1870 in Fuchs (1870a), Valvata simplex Fuchs, 1870 in Fuchs (1870b), and two new species (see discussion in Neubauer et al. 2014 for the rather confusing systematics applied in Bandel 2010). We follow Bandel and place the species within Muellerpalia .

The following subspecies have been introduced or ranked within Valvata simplex Fuchs, 1870:

1) Valvata bicincta Fuchs, 1870 [erroneously "bicinata" on p. 536; from captions and description there is no doubt about the correct name] from Tihany: It was considered a subspecies of Valvata simplex by Lőrenthey (1906, p. 166), what was followed by Wenz (1928a, p. 2475) and Strausz (1942, p. 36). Bandel (2010, p. 103) treated it as separate species and combined it with the new genus Muellerpalia . Current status: Muellerpalia bicincta .

2) Valvata carinata Fuchs, 1870 (p. 536) from Tihany: It was considered as subspecies of Valvata simplex by Pană et al. (1981) and Pană (2003), but recombined with the new genus Muellerpalia by Bandel (2010, p. 104). It is, however, a primary homonym of Valvata carinata Sowerby, 1834 (replacement name is introduced below).

3) Valvata simplex oecsensis Soós, 1934 (p. 189) from Öcs: Schlickum (1978, p. 246) clearly separated this taxon from " Hauffenia simplex " and retained it in Valvata . Wenz and Edlauer (1942, p. 83) elevated it to species level, what was followed by Papp (1953, p. 109), and Harzhauser and Binder (2004, p. 10). In some of the mentioned publications the name was erroneously emended to "oecsensis"; the correct emendation following ICZN rules is "ocsensis", since it is not derived from a German expression ( ICZN 1999, Article 32.5.2.1). Current status: Valvata ocsensis .

4) Valvata octonaria Brusina, 1902 (pl. 13) from Tihany: It was also ranked as subspecies of Valvata simplex by Wenz (1928a, p. 2476). Since it was not referred to by Bandel (2010), its generic affiliation is uncertain. Current status (needs revision): Muellerpalia haszprunari octonaria .

5) Valvata simplex polycincta Lőrenthey, 1906 (p. 167) from Tihany: It was synonymized with Valvata simplex octonaria by Wenz (1928a, p. 2476). Current status: junior synonym of Muellerpalia haszprunari octonaria .

6) Valvata simplex unicincta Lőrenthey, 1906 (p. 165) from Tihany ( Fehérpart): The status of this taxon is doubtful. It was not mentioned by Wenz (1928a) or Bandel (2010). Given the similarity with simplex and bicincta stated by Lőrenthey, it might fall into the intraspecific variability of either of these species. Current status (needs revision): Muellerpalia haszprunari unicincta .