Tylodinus pseudocavicrus Luna-Cozar, 2014

Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, Zootaxa 3788 (1), pp. 1-63 : 19

5. Tylodinus pseudocavicrus Luna-Cozar , sp. nov.

( Figures 98c , 36–37 , 102 , 130 )

Diagnosis. Length male, 5.0– 5.2 mm, female, 5.5–5.9 mm. Width male, 2.2–2.3 mm, female 2.5–2.6 mm. Body 2.2x longer than wide. Head with patch of whitish to yellow or light brown scales on middle of vertex, surface with dense punctures, somewhat irregular; frons concave. Pronotum in lateral view moderately convex; in dorsal view with longitudinal carina near middle, flanks with upper half rugose, lower half with deep and long punctures. Elytra with basal margin sinuate ( Figure 4 ); surface with punctures moderate, deep; tubercles distribution as follows: I2 with elongate tubercle near base of declivity; I3 with four tubercles, elongate subbasal tubercle, round tubercle at about midlength, two small elongate tubercles on declivity, one near middle, other near declivity apex; I4 with two tubercles, elongate, small, between tubercles of I3, other tubercle high, rounded at base of declivity, a little beyond I2 (this slightly higher than others); I5 with three elongate tubercles, first subbasal, second near midlength, both tubercles more or less the same size and shape as those on I 3 in same position and third tubercle little forward of base of declivity, much smaller than previous ones and that of neighboring tubercles, I6 with two elongate tubercles, small one located between tubercles on I5, and second near elytral declivity, I7 with two tubercles, one subbasal and round, other near middle of elytra, elongate, very small; tubercle on I2 on elytral declivity slightly smaller than that on I 4 in the same position. Metasternal tooth present. Male with ventrite 1 concave; ventrite 5 convex in middle, female with two apical lateral impressions, occasionally ventrite 2 depressed medially. Hind tibia with outer margin excavate, inner margin curved in apical ¼. Male genitalia ( Fig.102 ) moderately stout in lateral view; length in ventral view as long as apodemes, apex truncate; apical process moderately large.

Geographic distribution. México (Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2400–2910 m.

Derivation of specific name. Name derived from combining the Greek word pseudo = false, and the species name cavicrus .

Material examined. Total 9 males, 6 females. Holotype male ( CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16°44'33''N, 92°41'38.4''W), 18.ix.1991, coll. R . S. Anderson. Paratypes: México: Chiapas, Huixtán , 15 km E San Cristóbal, elev. 2500 m (16°44'48.84''N, 92°29'25.3''W), 28.v.2008, coll. LLAMA (Wa-A-05-1-32) — 1 male ( CMNC); San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16°44'33''N, 92°41'38.4''W), 18.ix.1991, coll. R . S. Anderson — 2 females ( CMNC); Cerro Huitepec ca. 5 km W San Cristóbal, elev. 2760 m (16°44'33''N, 92°41'38.4''W), 14.ix.2008, coll. R . S. Anderson — 1 female ( CMNC); Huitepec, S. Cristóbal , elev. 2480 m (16°45'1.4''N, 92°40'57.7''W), 29.v.2001, coll. LLAMA (Wa-A-05-2-32) — 1 male ( CMNC); Los Alcanfores, elev. 2450 m (16°44'12.47''N, 92°40'46.93''W), 25.iv.2006, coll. M. Girón — 1 male ( ECOS); same locality, 6.vii.2006, coll. J. L. Cozar — 1 male ( ECOS); Reserva Huitepec , elev. 2500 m (16°44'42.7''N, 92°40'52''W), 4.viii.1994, coll. B. Gomez — 1 male ( ECOS); same locality, 9.vii.2007, coll. M. G. Branstetter — 1 male ( CMNC); same locality, 17.ix.1991, coll. R . S. Anderson — 1 female ( CMNC); San Juan Chamula, Cerro Tzontehuitz, elev. 2800 m (16°49'10.2''N, 92°35'3.8''W), 15.vii.1994, coll. R . Jones — 1 female ( ECOS); Tzontehuitz ( Pico ) ca. 10 km NE San Cristóbal, elev. 2910 m (16°49'10.2''N, 92°35'3.8''W), 16.ix.1991, coll. R . S. Anderson — 1 female ( CMNC); Cerro Tzontehuitz , elev. 2864 m (16°48'40.8''N, 92°35'2.4''W), 29.vii.2005, coll. R . S. Anderson — 1 male ( CMNC); Cerro Tzontehuitz , elev. 2677 m (16°48'47''N, 92°34'24.8''W), 29.vii.2005, coll. R . S. Anderson — 1 male ( CMNC) .

Figures

FIGURES 92–99. 92–99. Dorsal and lateral habitus: 92–93. Tylodinus gibbosus, 94–95. Tylodinus kissingeri, 96–97. Tylodinus pinguis. 98. Median lobe apex of: a) Tylodinus elongatus, b) Tylodinus exiguus, c) Tylodinus pseudocavicrus, d) Tylodinus rufus, e) Tylodinus rufus (squashed), 99. Tylodinus zilchi, ventral and lateral habitus of median lobe.

FIGURES 36–43. Dorsal and lateral habitus: 36–37. Tylodinus pseudocavicrus, 38–39. Tylodinus rugosus. 40–41. Tylodinus buchanani. 42–43. Tylodinus exiguus.

FIGURES 100–109. Ventral and lateral habitus of median lobe: 100. Tylodinus andersoni, 101. Tylodinus cavicrus, 102. Tylodinus pseudocavicrus, 103. Tylodinus rugosus, 104. Tylodinus exiguus, 105. Tylodinus ixchel, 106. Tylodinus jonesi, 107. Tylodinus variabilis, 108). Tylodinus leoncortesi, 109. Tylodinus triumforium.

FIGURES 130–131. Map showing collecting sites. 130. Occurrences in México (Chiapas) and Central America of: Tylodinus zilchi (El Salvador, Honduras); Tylodinus andersoni (México), Tylodinus canaliculatus (México, Guatemala), Tylodinus triumforium (México), Tylodinus sepulturaensis (México), Tylodinus cavicrus (Guatemala), Tylodinus pseudocavicrus (México); 131. Occurrences in Chiapas, México of: Tylodinus complicatus, Tylodinus ixchel, Tylodinus dominicus, Tylodinus immundus, Tylodinus porvenirensis, Tylodinus branstetteri.

FIGURES 1–11. 1–5. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 6–11. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 3–5 in same plane as ventrites 1–2. 7. Tylodinus variabilis, male, note ventrites 3–5 not in same plane as ventrites 1–2. 8. Tylodinus coapillensis, female, note ventrites 1–2 strongly convex, ventrites 3–5 not in same plane as ventrites 1–2. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic).