Tozerium filzmoosense Cifer, 2020

Cifer, Tim, Goričan, Špela, Gawlick, Hans-Jürgen & Auer, Matthias, 2020, Pliensbachian, Early Jurassic radiolarians from Mount Rettenstein in the Northern Calcareous Alps, Austria, Acta Palaeontologica Polonica 65 (1), pp. 167-207 : 171-172

Tozerium filzmoosense Cifer sp. nov.

Fig. 6G–J .

ZooBank LCID: urn:lsid:zoobank.org:act:D07845CD-7397-456F-A9E5-9C0075F0C3CD

Etymology: Named after the town Filzmoos, which is located near Mount Rettenstein, Austria.

Type material: Holotype, PMS 2398 , sample Rö416: 170717 ( Fig. 6G ) . Paratypes: PMS 2397 , sample Rö 416: 170503 ; PMS 2398 , sample Rö 416: 170739 ; PMS 2393 , sample Rö 37: 171105; all from type locality. Type locality: Mount Rettenstein, Northern Calcareous Alps, Austria .

Type horizon: Sample Rö 416, grey marly limestone, Lower Pliensbachian .

Material.—Sample Rö37: stubs Rö37_3 (one specimen), Rö37_6 (two specimens); sample Rö416: stubs Rö416_5 three specimens), Rö416_7 (14 specimens); sample Rö417 one specimen), Mount Rettenstein , Northern Calcareous Alps, Austria, Lower Pliensbachian .

Diagnosis.— Tozerium with thick, three-bladed spines, throughout the whole length of the spine.

Description.—Cortical shell subspherical. Four spines in tetrahedral position. Pore frames irregularly shaped. Weak to relatively strong nodes are formed at pore frame vertices. Spines are three-bladed, tapering distally. Larger pores apparent at the base of the spines.

Dimensions.—See Table 1. Remarks.— Tozerium filzmoosense sp. nov. differs from Tozerium nascens Whalen and Carter in Carter et al., 1998 by having three-bladed spines. A similar species was described from the Hettangian ( Tozerium ? sp. B in Bertinelli and Marcucci 2011: 411, pl. 2: 16), but it differs from Tozerium filzmoosense sp. nov., by having the distal part of the spines circular in cross section. Tozerium filzmoosense sp. nov. may represent the advanced stage in the Tozerium lineage and is the youngest formally described species of the Tozerium which was considered to last appear at the end of the Sinemurian ( O’Dogherty et al. 2009). More recently, a Tozerium species with three-bladed but much thinner spines was reported from the Bajocian of east-central Oregon ( Tozerium sp. A in Yeh 2011: 6, pl. 12: 5, 16).

Stratigraphic and geographic range.—LowerPliensbachian. Northern Calcareous Alps ( Austria).

Figures

Fig. 6. Radiolarians from Mount Rettenstein, Austria, Pliensbachian, Early Jurassic. A. Empirea sp. 1, sample Rö416. B. Acaeniotylopsis ghostensis (Carter in Carter et al., 1988); sample Rö416. C, D. Liassobetraccium bavaricum (Kozur and Mostler, 1990); sample Rö37. E, F. Liassobetraccium verticispinosum (Kozur and Mostler, 1990); sample Rö37. G–J. Tozerium filzmoosense Cifer sp. nov. G. Holotype, sample Rö416: 170717. H. Paratype, sample Rö416: 170503. I. Paratype, sample Rö416: 170739. J. Paratype, sample Rö37: 171105. K–O. Loupanus pliensbachicus Cifer sp. nov. K. Holotype, sample Rö416: 170562. L. Paratype, sample Rö416: 170559. M. Paratype, sample Rö416: 170757. N. Paratype, sample Rö416: 170704. O. Paratype, sample Rö416: 170560. P, Q. Loupanus sp. 1; sample Rö417. R, S. Thurstonia? timberensis Whalen and Carter in Carter et al., 1998; sample Rö416. T. Thurstonia? minutaglobus Whalen and Carter in Carter et al., 1998; sample Rö416. U–W. Thurstonia? robusta Cifer sp. nov. U. Holotype, sample Rö417: Rö417_093.V. Paratype, sample Rö417: Rö417_090. W. Paratype, sample Rö417: Rö417_098.