Amphineurus (Rhamphoneurus) theischingeri, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.25. Amphineurus (Rhamphoneurus) theischingeri sp. nov.

Material examined.

Holotype: ♂, Chile, Osorno , Pucotrihue [40°32′S 73°42′W], 10/ 12-IV-1968, Peña (USNM) *. Paratypes: 2 ♂ and 1 ♀, Chile, Osorno , Bahia Mansa [40°35′S 73°44′W], South of Pucotrihue, 12-IV-1968, Peña (USNM) .

Etymology.

The species is named after Gunther Theischinger, an expert on Tipulomorpha who revised the Australian Amphineurus .

Diagnosis.

This species is distinguished by an absence of markings near R2, r-m as long as basal deflection of R5, and cross-vein m-cu straight and inserted at bM. Furthermore, this species is characterized by male terminalia with curved sheath of aedeagus, straight medial branch of clasper of gonostylus and mesal lobes with slender, vertical tips. The thorax is also distinguishable by narrow knob of halter and katepisternum as long as meron.

Description.

Wing length 6.30 mm, width 1.77 mm. - Coloration: General coloration dark brown. Rostrum and palpus brownish black. First segment of antenna yellow, remainding segments dark yellow. Head brownish gray. Eye black. Thorax dark-brown, pleura light brown. Scutum without stripes. Halter brownish-yellow. Coxae brownish-yellow, legs dark brown. Abdomen brownish black. - Head (dorsal view Fig. 30B , ventral view Fig. 30A ): Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel; terminal flagellomere longer than penultimate. - Thorax (lateral view Fig. 30E , dorsal view Fig. 30F ): Anatergite shorter than katatergite. Halter with narrow knob. Wing (Fig. 30D ) clear of distinctive markings; veins near fork of bM (Fig. 30C ) visible; M1+2 longer than basal deflection of M1; R2 faded; R2+3+4 longer than R2+3. - Female terminalia (Fig. 30H ): Female tergite IX shorter and wider than tergite X; hypogynial valve very slender; cercus longer than hypogynial valve, thick, and slightly curved dorsad. - Male terminalia (Fig. 30G ): Male tergite IX with V-shaped median notch. Distal tip of ventral branch of gonocoxite expanded, setose, with slight curve ventrally. Dorsal branch of gonocoxite shorter than half-length of ventral branch. Lobe of gonostylus with rounded lobule; lobule longer than wide, more than half as long as lobe of gonostylus, wider than stem. Clasper of gonostylus with similarly shaped branches: lateral branch rounded; medial branch conical. Mesal lobes of gonocoxite asymmetrical; both with long, narrow, straight distal projection; right mesal lobe wider. Sheath of the aedeagus curved halfway along length, darkened only on tip.

Remarks.

Specimens of this new species were previously identified by C.P. Alexander as A. (R.) nullus Alexander. This species resembles A. (R.) billighami sp. nov. but differs mainly in the attachment of m-cu, setae concentration in gonocoxite, shape of male tergite IX and mesal lobe.

Figures

Figure 30. Amphineurus (Rhamphoneurus) theischingeri sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX.