Eusphalerum obscuriceps, Motschulsky, 1860

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

obscuriceps Motschulsky, 1860 , see: angusticollis Mannerheim, 1830 .

omalinus Zetterstedt, 1828: 46 ( Anthophagus ; Type locality: Lapponia Norvegica. Nordlandia: Bjorkvik; Evenas; Lingen. Suecia: Johannis Ro. Insula Dyroe : Hamnvigen. Insula Senjen: Goebostad). — C. Sahlberg, 1827: 274 ( Anthophagus ; [Note: Misidentification: Bernhauer and Schubert, 1910: 80 cited alpinus Sahlberg as a synonym of omalinus Zetterstedt ; Sahlberg did not describe the species as new, he attributed it to Fabricius]; characters; Finland). — Zetterstedt, 1838: 48 ( Anthophagus ; characters; habitat; Lapland). — Erichson, 1840: 849 ( Anthophagus ; characters; Lapland). — Heer, 1841: 573 ( Anthophagus ; characters). — Kiesenwetter, 1845: 23 ( Anthophagus ; notes). — Kiesenwetter, 1847a: 54 ( Anthophagus ; characters; Germany; Lapland; Sweden; Switzerland). — Märkel and Kiesenwetter, 1848a: 327 ( Anthophagus ; notes; Carpathian Alps). — Redtenbacher, 1849: 746 ( Anthophagus ; characters; Austria). — Fairmaire and Laboulbène, 1856: 628 ( Anthophagus ; characters; France). — Redtenbacher, 1857: 242 ( Anthophagus ; characters; Austria). — Kraatz, 1857d: 921 ( Anthophagus ; characters; Germany). — Thomson, 1861: 179 ( Anthophagus ; characters; Scandinavia). — Hochhuth, 1862: 102 ( Anthophagus ; Russia). — Fauvel, 1865a: 315 [= 1865b: 65] ( Anthophagus ; habitat; France). — Redtenbacher, 1874: 264 ( Anthophagus ; characters; Austria). — Seidlitz, 1875: 242 ( Anthophagus ; characters; Baltic region). — Mulsant and Rey, 1880: 40 ( Anthophagus ; characters; France). — Ganglbauer, 1895: 709 ( Anthophagus ; characters; north Europe; Sudeten; Alps). — Luze, 1902d: 523 ( Anthophagus ; characters; Sudeten; northern Europe; Alps). — Reitter, 1909: 183 ( Anthophagus ; characters; Germany). — Bernhauer and Schubert, 1910: 80 ( Anthophagus ; catalog). — Portevin, 1929: 435 ( Anthophagus ; characters; France). — Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog). — C. Koch, 1933a: 142, 143 ( Anthophagus ; subgenus Dimorphoschelus ; figure). — C. Koch, 1934a: 223 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Lapland; Finland; Sweden; Norway; Russia; Germany; Czechoslovakia; Austria; Croatia; Italy; Switzerland; France). — Palm, 1948: 125 ( Anthophagus ; characters; Sweden; Norway; Finland). — Horion, 1963: 147 ( Anthophagus ; subgenus Dimorphoschelus ; Norway; Sweden; Finland; Russia; Siberia; Czech Republic; Slovakia; Romania; Slovenia; Croatia; Germany; Austria). — Tóth, 1982: 97 ( Anthophagus ; characters). — Zanetti, 1987: 416 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Italy). — Lucht, 1987: 90 ( Anthophagus ; distribution of nominate subspecies; checklist; Sweden). — Zanetti, 1995: 9 ( Anthophagus ; Italy). — Cibuļskis, 1999: 36 ( Anthophagus ; list; Latvia). - DISTRIBUTION: Europe, Russia, Caucasus.

arpedinus Hochhuth, 1849: 201 ( Anthophagus ; Type locality: Kaukasus). — Luze, 1902d: 524 ( Anthophagus ; characters; Caucasus). — Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog). — C. Koch, 1933a: 142 ( Anthophagus ; subspecies of omalinus ). — C. Koch, 1934a: 225 ( Anthophagus ; synonym of omalinus ). — Tikhomirova, 1973a: 141 ( Anthophagus ; subgenus Dimorphoschelus ; checklist; Caucasus).

ruficeps Motschulsky, 1860: 547 ( Heterops ; Type locality: Sibérie orientale au delà du lac Baical ; [Note: See lectotype designation by Hammond, 1970]). — Heyden, 1880: 82 ( Heterops ; Siberian region). — Bernhauer and Schubert, 1910: 70 ( Amphichroum ; catalog). — Hammond, 1970: 69, 70 ( Anthophagus ; lectotype designation; notes; synonym of omalinus ).

homalinus Fauvel, 1871 a: 149 [= 1872: 123] ( Anthophagus ; emendation of omalinus ). — J. Sahlberg, 1876: 207 ( Anthophagus ; Finland). — Luze, 1902d: 523 ( Anthophagus ; synonym of omalinus ). — Bernhauer and Schubert, 1910: 80 ( Anthophagus ; synonym of omalinus ). — C. Koch, 1933a: 142 ( Anthophagus ; synonym of omalinus ). — Zanetti, 1987: 416 ( Anthophagus ; synonym of omalinus ).

arrowi C. Koch, 1933: 29 ( Anthophagus ; subspecies of omalinus ; Type locality: Europa centralis). — C. Koch, 1933a: 142 ( Anthophagus ; subspecies of omalinus ). — C. Koch, 1934a: 223 ( Anthophagus ; subgenus Dimorphoschelus ; race of omalinus ; characters). — Lohse, 1964: 67 ( Anthophagus ; subgenus Dimorphoschelus ; subspecies of omalinus ; characters; central Europe). — Zanetti, 1987: 416 ( Anthophagus ; subgenus Dimorphoschelus ; subspecies of omalinus ; characters; Italy). — Lucht, 1987: 90 ( Anthophagus ; subspecies of omalinus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland). — Boháč, 1993: 42 ( Anthophagus ; subgenus Anthophagus ; subspecies of omalinus ; Slovakia; Czech Republic). — Zanetti, 1995: 9 ( Anthophagus ; subspecies of omalinus ; Italy).

ursus Havelka and R. Dvořák, 1953: 28 ( Anthophagus ; aberration of omalinus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Carniola: Medvedi Dolina, Vallée d’Ours).

palustris Heer, 1841 , see: caraboides Linné, 1758 .

paratus Havelka and R. Dvořák, 1953 [Note: Nomen nudum], see: alpinus Fabricius, 1793 .

picipennis Gredler, 1863 , see: spectabilis Heer, 1839 .

pollinensis C. Koch, 1934 , see: torretassoi C. Koch, 1933 .

praeustus P. W. J. Müller, 1821: 228 ( Anthophagus ; Type locality: Odenbach). — Runde, 1835: 23 ( Lesteva ; characters; Germany). — Heer, 1839: 194 ( Anthophagus ; characters; Switzerland). — Erichson, 1840: 852 ( Anthophagus ; characters; Germany; France). — Redtenbacher, 1849: 747 ( Anthophagus ; characters; Austria). — Fairmaire and Laboulbène, 1856: 629 ( Anthophagus ; characters; France). — Redtenbacher, 1857: 243 ( Anthophagus ; characters; Austria). — Kraatz, 1857d: 924 ( Anthophagus ; characters; Germany). — Fauvel, 1871a: 141 [= 1872: 115] ( Anthophagus ; characters; France; Germany; Switzerland; Italy; Austria). — Fauvel, 1873b: 119 [= 1873c: 12] ( Anthophagus ; Sardinia). — Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria). — Mulsant and Rey, 1880: 48 ( Anthophagus ; subgenus Phaganthus ; characters; France). — Ganglbauer, 1895: 709 ( Anthophagus ; subgenus Phaganthus ; characters; middle Europe). — Luze, 1902d: 527 ( Anthophagus ; characters; central Europe). — Reitter, 1909: 183 ( Anthophagus ; subgenus Phaganthus ; characters; Germany). — Bernhauer and Schubert, 1910: 80 ( Anthophagus ; cited as pracustus; catalog). — Portevin, 1929: 435 ( Anthophagus ; characters; France). — Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog). — C. Koch, 1933a: 141, 143 ( Anthophagus ; subgenus Phaganthus ; figure). — C. Koch, 1934a: 158 ( Anthophagus ; subgenus Phaganthus ; characters; Germany; France; Switzerland; Portugal). — Horion, 1963: 138 ( Anthophagus ; subgenus Phaganthus ; Holland; Belgium; France; Spain; Portugal; Switzerland; Sardinia; Italy; Romania; Czech Republic; Germany; Austria). — Lohse, 1964: 64 ( Anthophagus ; subgenus Phaganthus ; characters; central Europe). — Smetana, 1964c: 164 ( Anthophagus ; subgenus Phaganthus ; Czechoslovakia). — Tóth, 1982: 92 ( Anthophagus ; characters; Hungary). — Zanetti, 1987: 393 ( Anthophagus ; species not in Italy).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Boháč, 1993: 41 ( Anthophagus ; subgenus Phaganthus ; Czech Republic).

— Zerche, 2000: 52 ( Anthophagus ; discussion; nomen protectum).

- DISTRIBUTION: Europe.

nigrophthalmus Block, 1799: 117 ( Staphylinus ; Type locality: Plauischer Grund bei Dresden).

— Gemminger and Harold, 1868: 658 ( Anthophagus ; synonym of plagiatus ).

— Fauvel, 1869: 493 ( Anthophagus ; synonym of plagiatus ).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; synonym of plagiatus ).

— Zerche, 2000: 52 ( Anthophagus ; synonym of praeustus ; nomen protectum).

bimaculatus Lacordaire, 1835: 481 ( Lesteva ; Type locality: Fontainbleau).

— Erichson, 1840: 852 ( Anthophagus ; synonym of praeustus ).

— Redtenbacher, 1849: 747 ( Anthophagus ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 629 ( Anthophagus ; synonym of praeustus ).

— Kraatz, 1857d: 925 ( Anthophagus ; synonym of praeustus ).

— Mulsant and Rey, 1880: 49 ( Anthophagus ; synonym of praeustus ).

— Ganglbauer, 1895: 709 ( Anthophagus ; synonym of praeustus ).

— Luze, 1902d: 527 ( Anthophagus ; synonym of praeustus ).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; synonym of praeustus ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of praeustus ).

pseudosudeticus C. Koch, 1934 [Note: Nomen nudum], see: alpinus Fabricius, 1793 .

puberulum Märkel and Kiesenwetter, 1848 , see: Eusphalerum .

puncticollis Weise, 1875 , see: Geodromicus .

pyrenaeus Brisout, 1863 , see: alpinus Fabricius, 1793 .

rotundicollis Heer, 1839: 195 ( Anthophagus ; Type locality: Frugmatt).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— Mulsant and Rey, 1880: 51 ( Anthophagus ; subgenus Phaganthus ; characters; France).

— Ganglbauer, 1895: 709 ( Anthophagus ; subgenus Phaganthus ; characters; Alps).

— Luze, 1902d: 528 ( Anthophagus ; characters; East Tirol).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141, 143 ( Anthophagus ; subgenus Phaganthus ; figure).

— C. Koch, 1934a: 160 ( Anthophagus ; subgenus Phaganthus ; characters; Alpes; Austria; Transsylvania; Alps).

— Horion, 1935: 169 ( Anthophagus ; subgenus Phaganthus ; characters; Alps; Austria; Germany).

— Horion, 1963: 140 ( Anthophagus ; subgenus Phaganthus ; France; Switzerland; Italy; Germany; Austria; Romania).

— Lohse, 1964: 64 ( Anthophagus ; subgenus Phaganthus ; characters; central Europe).

— Tóth, 1982: 92 ( Anthophagus ; characters).

— Zanetti, 1987: 392 ( Anthophagus ; subgenus Phaganthus ; characters; Italy).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Austria).

- DISTRIBUTION: Germany, France, Switzerland Italy, Austria.

brevicornis Kiesenwetter, 1861: 376 ( Anthophagus ; Type locality: Saas am Ufer des Vispbaches; Macugnaga am Ufer der Anza; Annivier-Thal an der Nanvisanche).

— Fauvel, 1871a: 139 [= 1872: 113] ( Anthophagus ; characters; France; Germany; Switzerland; Italy; Austria).

— Fauvel, 1874: 324 [= 1874b: 30] ( Anthophagus ; synonym of brevicornis ).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— Mulsant and Rey, 1880: 51 ( Anthophagus ; synonym of rotundicollis ).

— Ganglbauer, 1895: 709 ( Anthophagus ; synonym of rotundicollis ).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; synonym of rotundicollis ).

— Portevin, 1929: 435 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; synonym of rotundicollis ).

— C. Koch, 1934a: 160 ( Anthophagus ; subgenus Phaganthus ; aberration of rotundicollis ).

— Tóth, 1982: 92 ( Anthophagus ; aberration of rotundicollis ; characters).

— Zanetti, 1987: 392 ( Anthophagus ; synonym of rotundicollis ).

crassicornis Mulsant and Rey, 1861: 179 [= 1862: 163] ( Anthophagus ; Type locality: Chamouni).

— Fauvel, 1871a: 139 [= 1872: 113] ( Anthophagus ; synonym of brevicornis ).

— Mulsant and Rey, 1880: 51 ( Anthophagus ; synonym of rotundicollis ).

— Ganglbauer, 1895: 709 ( Anthophagus ; synonym of rotundicollis ).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; synonym of rotundicollis ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of rotundicollis ).

imitator Luze, 1902d: 528 ( Anthophagus ; aberration of rotundicollis ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; no locality cited).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; aberration of rotundicollis ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of brevicornis ).

roubali C. Koch, 1934 , see: angusticollis Mannerheim, 1830 .

ruficeps Motschulsky, 1860 , see: omalinus Zetterstedt, 1828 .

schatzmayri C. Koch, 1933: 30 ( Anthophagus ; Type locality: Brussa).

— C. Koch, 1933a: 142 ( Anthophagus ; subgenus Dimorphoschelus ).

— C. Koch, 1934a: 237 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Asia Minor).

— Tikhomirova, 1973a: 141 ( Anthophagus ; subgenus Dimorphoschelus ; checklist; Asia Minor). - DISTRIBUTION: Turkey.

scheerpeltzi C. Koch, 1933 , see: muticus Kiesenwetter, 1850 .

schneideri Eppelsheim, 1878: 125 ( Anthophagus ; Type locality: Elisabetpol).

— Luze, 1902d: 517 ( Anthophagus ; characters; Caucasus; Transcaucasian region).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— C. Koch, 1933a: 142 ( Anthophagus ; subgenus Dimorphoschelus ).

— C. Koch, 1934a: 221 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Caucasus).

- DISTRIBUTION: Armenia, Azerbaijan, Georgia.

scutellaris Erichson, 1840: 851 ( Anthophagus ; Type locality: Genevae).

— Heer, 1841: 572 ( Anthophagus ; [Note: Misidentification: Bernhauer and Schubert, 1910: 81 cited caraboides variety b Heer as a synonym of scutellaris ; the variety is unavailable]; characters). — Heer, 1841: 572 ( Anthophagus ; characters; Switzerland).

— Kiesenwetter, 1845: 23 ( Anthophagus ; notes).

— Kiesenwetter, 1847a: 58 ( Anthophagus ; characters; Switzerland).

— Fairmaire and Laboulbène, 1856: 628 ( Anthophagus ; characters; France).

— Kraatz, 1857d: 916 ( Anthophagus ; notes).

— Fauvel, 1865a: 314 [= 1865b: 64] ( Anthophagus ; habitat; France).

— Fauvel, 1871a: 142 [= 1872: 116] ( Anthophagus ; characters; France; Switzerland; Italy).

— Mulsant and Rey, 1880: 43 ( Anthophagus ; subgenus Phaganthus ; characters; France).

— Ganglbauer, 1895: 709 ( Anthophagus ; subgenus Phaganthus ; characters; western Alps; Jura). — Luze, 1902d: 525 ( Anthophagus ; characters; Jura; France; Piemont; Swiss and Tirolian Alps; Austria).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— Portevin, 1929: 435 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141, 143 ( Anthophagus ; subgenus Phaganthus ; figure).

— C. Koch, 1934a: 164 ( Anthophagus ; subgenus Phaganthus ; characters; Europe).

— Horion, 1935: 169 ( Anthophagus ; subgenus Phaganthus ; characters; Alps; Austria).

— Horion, 1963: 140 ( Anthophagus ; subgenus Phaganthus ; France; Italy; Switzerland; Germany; Austria).

— Lohse, 1964: 64 ( Anthophagus ; subgenus Phaganthus ; characters; central Europe).

— Tóth, 1982: 91 ( Anthophagus ; characters).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Austria; Switzerland).

— Zanetti, 1995: 9 ( Anthophagus ; Italy).

- DISTRIBUTION: Germany, France, Switzerland, Austria, Italy.

sinuatus Sharp, 1889 , see: Geodromicus .

solarii C. Koch, 1934 , see: bicornis Block, 1799 .

spectabilis Heer, 1839: 197 ( Anthophagus ; Type locality: in Valesiae alpibus).

— Kiesenwetter, 1845: 23 ( Anthophagus ; notes).

— Fairmaire and Laboulbène, 1856: 627 ( Anthophagus ; characters; France).

— Redtenbacher, 1857: 242 ( Anthophagus ; characters; Austria).

— Kraatz, 1857d: 914 ( Anthophagus ; characters; Austria).

— Fauvel, 1865a: 314 [= 1865b: 64] ( Anthophagus ; France).

— Fauvel, 1871a: 144 [= 1872: 118] ( Anthophagus ; characters; France; Switzerland; Germany). — Redtenbacher, 1874: 264 ( Anthophagus ; characters; Austria).

— Mulsant and Rey, 1880: 24 ( Anthophagus ; characters; France).

— Ganglbauer, 1895: 707 ( Anthophagus ; characters; Alps; Vosges).

— Luze, 1902d: 511 ( Anthophagus ; characters; Alps; Voges).

— Reitter, 1909: 182 ( Anthophagus ; characters; Germany).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— Portevin, 1929: 434 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141 ( Anthophagus ; subgenus Anthophagus ).

— C. Koch, 1934a: 173 ( Anthophagus ; subgenus Anthophagus ; characters; Switzerland; France; Italy).

— Horion, 1963: 142 ( Anthophagus ; subgenus Anthophagus ; France; Italy; Switzerland; Germany; Austria).

— Lohse, 1964: 64 ( Anthophagus ; subgenus Anthophagus ; characters; central Europe).

— Zanetti, 1987: 401 ( Anthophagus ; subgenus Anthophagus ; characters; Italy).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Austria; Switzerland; France).

— Zanetti, 1995: 9 ( Anthophagus ; Italy).

- DISTRIBUTION: France, Switzerland, Germany, Austria, Italy.

picipennis Gredler, 1863 a: 166 ( Anthophagus ; Type locality: Not cited, collected in the Alps ).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— Fauvel, 1876a: 54 [= 1876: 239] ( Anthophagus ; synonym of picipennis ).

— Luze, 1902d: 511 ( Anthophagus ; synonym of spectabilis ).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; synonym of spectabilis ).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; synonym of spectabilis ).

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of spectabilis ).

— C. Koch, 1934a: 173 ( Anthophagus ; subgenus Anthophagus ; aberration of spectabilis ). muelleri Bernhauer, 1901: 539 ( Anthophagus ; variety of spectabilis ; Type locality: Bregenz auf den Vorarlberger Alpen).

— Luze, 1902d: 511 ( Anthophagus ; variety of spectabilis ).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; variety of spectabilis ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of picipennis ).

— C. Koch, 1934a: 176 ( Anthophagus ; subgenus Anthophagus ; aberration of spectabilis ; characters).

— Zanetti, 1987: 401 ( Anthophagus ; synonym of spectabilis ).

speculicollis C. Koch, 1933 , see: alpinus Fabricius, 1793 .

subfasciatus Luze, 1902 [Note: Nomen nudum], see: bicornis Block, 1799 .

subtilis Sharp, 1889 , see: Liophilydrodes .

sudeticus Kiesenwetter, 1846: 22 ( Anthophagus ; Type locality: Riesengebirge ; Spieglitzer Schneeberg im Glatzer Gebirge).

— Redtenbacher, 1849: 828 ( Anthophagus ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 626 ( Anthophagus ; characters; France).

— Redtenbacher, 1857: 243 ( Anthophagus ; characters; Austria).

— Kraatz, 1857d: 920 ( Anthophagus ; characters; Germany).

— Fauvel, 1871a: 147 [= 1872: 121] ( Anthophagus ; characters; Italy; Austria).

— Redtenbacher, 1874: 264 ( Anthophagus ; characters; Austria).

— Mulsant and Rey, 1880: 16 ( Anthophagus ; characters; France).

— Ganglbauer, 1895: 707 ( Anthophagus ; characters; Sudeten).

— Luze, 1902d: 520 ( Anthophagus ; characters; Sudeten; Carpathian and Transylvanian Alps).

— Reitter, 1909: 182 ( Anthophagus ; characters; Germany).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog).

— C. Koch, 1933a: 142, 143 ( Anthophagus ; subgenus Anthophagus ; figure).

— C. Koch, 1934a: 205 ( Anthophagus ; subgenus Anthophagus ; characters; central Europe).

— Horion, 1963: 144 ( Anthophagus ; subgenus Anthophagus ; Czech Republic; Slovakia; Romania; Germany; Austria).

— Lohse, 1964: 66 ( Anthophagus ; subgenus Anthophagus ; characters; central Europe).

— Tóth, 1982: 94 ( Anthophagus ; characters).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Poland; Czechoslovakia).

— Boháč, 1993: 42 ( Anthophagus ; subgenus Dimorphoschelus ; Slovakia; Czech Republic).

- DISTRIBUTION: Poland, Germany, Austria, Italy, Czech Republic, Slovakia, Romania, Carpathians.

testaceus Gravenhorst, 1802 , see: caraboides Linné, 1758 .

torretassoi C. Koch, 1933: 27 ( Anthophagus ; Type locality: Appennino abruzzese, Lazio).

— C. Koch, 1933a: 142 ( Anthophagus ; subgenus Anthophagus ).

— C. Koch, 1934a: 185 ( Anthophagus ; subgenus Anthophagus ; characters; Italy).

— Zanetti, 1987: 405 ( Anthophagus ; subgenus Anthophagus ; characters; Italy).

— Zanetti, 1995: 10 ( Anthophagus ; Italy).

- DISTRIBUTION: Italy.

pollinensis C. Koch, 1934 a: 185, 188 ( Anthophagus ; subgenus Anthophagus ; race of torretassoi ; Type locality: Lucania, Mte. Pollino, Duglia).

— Zanetti, 1987: 405 ( Anthophagus ; subgenus Anthophagus ; subspecies of torretassoi ; characters; Italy).

— Zanetti, 1995: 10 ( Anthophagus ; subspecies of torretassoi ; Italy).

transversus Motschulsky, 1857 a: 494 ( Anthophagus ; Type locality: Alpes de la Carniole).

— Fauvel, 1865c: 18 [= 1866: 25] ( Anthophagus ; synonym of austriacus ).

— Kraatz, 1867: 415 ( Anthophagus ; synonym of austriacus ).

— Fauvel, 1871a: 141 [= 1872: 115] ( Anthophagus ; synonym of alpestris ).

— Ganglbauer, 1895: 708 ( Anthophagus ; synonym of alpestris ).

— Luze, 1902d: 512 ( Anthophagus ; synonym of alpestris ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; aberration of alpestris ).

— C. Koch, 1933a: 142 ( Anthophagus ; subspecies of dinaricus ).

— C. Koch, 1934a: 234 ( Anthophagus ; subgenus Dimorphoschelus ; race of dinaricus ; characters).

— Zanetti, 1986: 96 ( Anthophagus ; cited as valid species).

— Zanetti, 1987: 419 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Italy).

— Zanetti, 1995: 10 ( Anthophagus ; Italy).

- DISTRIBUTION: Italy, Austria, Bosnia Hercegovina, Slovenia.

dinaricus Apfelbeck, 1909: 501 ( Anthophagus ; Type locality: Bosnia sept. occ.: in monte Klekovača ).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog).

— C. Koch, 1933: 30 ( Anthophagus ; Italy).

— C. Koch, 1933a: 142, 143 ( Anthophagus ; subgenus Dimorphoschelus ; figure).

— C. Koch, 1934a: 231 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Herzegowina;

— Horion, 1963: 148 ( Anthophagus ; subgenus Dimorphoschelus ; Austria; Bosnia Hercegovina; Slovenia).

— Lohse, 1964: 67 ( Anthophagus ; subgenus Dimorphoschelus ; characters; central Europe).

— Zanetti, 1986: 96 ( Anthophagus ; subspecies of transversus ).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Austria).

— Zanetti, 1987: 419 ( Anthophagus ; subgenus Dimorphoschelus ; subspecies of transversus ; characters; Italy).

— Lohse and Lucht, 1989: 127 ( Anthophagus ; subspecies of transversus ).

— Zanetti, 1995: 10 ( Anthophagus ; subspecies of transversus ; Italy).

trimaculatus Luze, 1902 , see: caraboides Linné, 1758 .

ursus Havelka and R. Dvořák, 1953 [Note: Nomen nudum], see: omalinus Zetterstedt, 1828 .

variegatus Gistel, 1857: 5 ( Anthophagus ; [Note: Nomen dubium]; Type locality: Styria).

- DISTRIBUTION: Austria.

vehemans Gistel, 1857: 85 ( Anthophagus ; [Note: Nomen dubium]; Type locality: Tyrolis).

- DISTRIBUTION: Austria.

verticalis Say, 1831 , see: Geodromicus .

villosus Waltl, 1838 , see: Lesteva .

Arpedium

[6 species; Nearctic and Palaearctic Region]

Arpedium Erichson, 1839 a: 618 (species included: quadrum ). TYPE SPECIES: Omalium quadrum Gravenhorst , fixed by monotypy.

— Erichson, 1840: 858 (characters).

— Redtenbacher, 1849: 748 (characters).

— Schaum, 1852: 30 (list of species; Europe).

— Lacordaire, 1854: 139 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 633 (characters).

— Jacquelin du Val, 1857: 72 (characters).

— Redtenbacher, 1857: 246 (characters).

— Kraatz, 1857d: 957 (characters).

— Thomson, 1858: 37 (characters).

— G. Waterhouse, 1858: 31 (catalog of British species).

— Thomson, 1859: 49 (characters; type species: quadrum ).

— Schaum, 1859: 31 (catalog; European species).

— Thomson, 1861: 188 (characters).

— Gredler, 1863: 123 (list of species from Tirol).

— Wencker and Silbermann, 1866: 36 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 662 (catalog).

— Jacquelin du Val, 1868: 80 (catalog; European species).

— Fauvel, 1871a: 111 [= 1872: 85] (characters; key to species).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 270 (characters).

— Fauvel, 1875a:VI [= 1875b: 208] (catalog).

— Fauvel, 1878d: 224 [= 1878e: 60] (key to North American species).

— Mulsant and Rey, 1880: 144 (characters; key to French species).

— Heyden, 1880: 82 (list of species of Siberian region).

— Duvivier, 1883: 196 (catalog).

— Fowler, 1888: 407 (notes).

— Seidlitz, 1891: 343 (key to some European species).

— Seidlitz, 1891a: 365 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 119 (list of species of Europe and Caucasus).

— Heyden, 1893: 48 (list of species of Siberian region).

— Ganglbauer, 1895: 717 (characters; key to middle European species).

— Heyden, Reitter, and Weise, 1906: 134 (list of species of Europe and Caucasus).

— Reitter, 1909: 186 (characters; key to German species).

— Bernhauer and Schubert, 1910: 67 (world catalog; 15 species).

— Blatchley, 1910: 477 (characters; key to Indiana species).

— Petri, 1912: 54 (list of species of Siebenbürgen).

— Johansen, 1914: 600 (characters; key to Danish species).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 333 (catalog for Palaearctic region).

— Cameron, 1925: 11 (catalog of Indian species).

— Porta, 1926: 26 (characters; key to species of Italy).

— Portevin, 1929: 440 (key to French genera).

— Cameron, 1930: 155 (characters).

— Roubal, 1930: 314 (catalog; Slovakia).

— Munster, 1933 (revision of and key to Norwegian species).

— Joy, 1932: 102 (key to British species).

— Scheerpeltz, 1933: 1060 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 109 (characters; key to species of Sweden).

— Tottenham, 1949: 357, 407 (type species: quadrum Gravenhorst ; list of British species).

— Blackwelder, 1952: 62 (type species: quadrum ).

— Tottenham, 1954: 31 (characters; British species).

— Hatch, 1957: 49, 53 (characters; key to species of Pacific Northwest).

— Horion, 1963: 117 (list of central European species).

— Székessy, 1963: 20 (characters).

— Lohse, 1964: 55 (characters; key to central European species).

— Moore, 1966: 48, 51 (characters; notes).

— Scheerpeltz, 1968a: 14 (catalog of Austrian species).

— Tikhomirova, 1973a: 140 (checklist of species of USSR).

— Moore and Legner, 1975: 184 (catalog; North America north of Mexico).

— Pope, 1977: 22 (list of British species).

— Topp, 1978: 308 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 61 (catalog; Poland).

— Moore and Legner, 1979: 180 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Uhlig, 1979: 243 (some species collected in Germany).

— Uhlig and Vogler, 1981: 87 (list of some species collected in Germany; habitat and collecting notes).

— Tóth, 1982: 81 (characters; habitus; key to species of Hungary).

— Campbell, 1984: 506 (characters; revision of and key to species of North America ).

— Ádám, 1985: 251 (list of species from southeast Hungary).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Zanetti, 1987: 307 (characters; habitus; species of Italy).

— Ádám, 1987: 137 ( Arpedium ; collecting notes; Hungary).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— Zanetti, 1995: 8 (list of species of Italy).

— M. Hansen, 1996: 95 (list of Danish species).

— Downie and Arnett, 1996: 431 (characters; key to species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 339 (3 Nearctic species; notes; characters in key).

alpinum Fauvel, 1871 , see: quadrum Gravenhorst, 1806 .

angulare Fauvel, 1878 , see: cribratum Fauvel, 1878 .

angulare Fauvel, 1878d: 226 [= 1878e: 62] ( Arpedium ; Type locality: Pointe des Pins, Sault St. Marie, Lake Superior ; [Note: See lectotype designation by Campbell, 1984]. Also cited from Detroit, Michigan).

— Campbell, 1984: 508 ( Arpedium ; lectotype designation: Pointe des Pins , Sault St. Marie, Lake

Superior, Ontario; synonym of cribratum ).

— Newton, Thayer, Ashe, and Chandler, 2000: 339 ( Arpedium ; appears to be distinct species).

- DISTRIBUTION: Canada; USA.

beringense Van Dyke, 1921, see: Eucnecosum .

brevicollis Mäklin, 1852 , see: Porrhodites .

brunnescens J. Sahlberg, 1871 , see: Eucnecosum .

columbiense Hatch, 1957 , see: cribratum Fauvel, 1878 .

creticum Scheerpeltz, 1958, see: Deliphrosoma .

cribratum Fauvel, 1878d: 224 [= 1878e: 60] ( Arpedium ; Type locality: Detroit , Michigan;

[Note: See lectotype designation by Campbell, 1984]. Originally cited from Michigan, Detroit ; Illinois).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; catalog).

— Blatchley, 1910: 478 ( Arpedium ; characters; Indiana).

— Legner and Moore, 1977: 175 ( Arpedium ; found under bark).

— Campbell, 1984: 508 ( Arpedium ; lectotype designation: Detroit, Michigan; characters; British

Columbia; New Brunswick; Ontario; Quebec; Illinois; Iowa; Maine; Massachusetts; Michigan;

Missouri; Nebraska; New Hampshire; New York; Ohio; Wisconsin).

— Campbell and Davies, 1991: 89 ( Arpedium ; checklist of species; Canada).

— Levesque and Levesque, 1996: 289 ( Arpedium ; seasonal abundance; Quebec).

— Downie and Arnett, 1996: 431 ( Arpedium ; characters; USA).

- DISTRIBUTION: Canada; USA.

columbiense Hatch, 1957: 53 ( Arpedium ; Type locality: Barkerville, B. C. Also cited from

Idaho).

— Steel, 1964a: 249 ( Arpedium ; synonym of quadrum ).

— Campbell, 1984: 508 ( Arpedium ; synonym of cribratum ).

engadinicum Heyden, 1863 , see: Eucnecosum .

gyllenhali C. Sahlberg, 1830 , see: Eucnecosum .

heydeni Heer, 1839 , see: quadrum Gravenhorst, 1806 .

humile Erichson, 1840 , see: Philorinum .

impressum Zetterstedt, 1828 , see: Eucnecosum .

kittilense Munster, 1933 , see: Eucnecosum .

libanicum Fauvel, 1875 , see: Deliphrosoma .

limbatum Gebler, 1848 , see: quadrum Gravenhorst, 1806 .

lindbergi Scheerpeltz, 1958 , see: Deliphrosoma .

maculicolle Mannerheim, 1843 , see: Amphichroum .

major Bernhauer, 1902 , see: Deliphrosoma .

mandli Scheerpeltz, 1937 , see: Deliphrosoma .

mixtum Bernhauer, 1902 , see: Eucnecosum .

morvani Jarrige, 1971 , see: Deliphrosoma .

myops Haliday, 1841 , see: Philorinum .

nepalicum Coiffait, 1982: 145 ( Arpedium ; Type locality: Nepal occidental: Dzunda, Khola

Tal, près Talphi, 3000–3500 m).

- DISTRIBUTION: Nepal.

norvegicum Munster, 1933 , see: Eucnecosum .

pallens Motschulsky, 1857 , see: Phloeonomus .