Eudyasmus Pascoe, 1885: 275

Setliff, Gregory, Pancini, Lorenzo & Bramanti, Andrea, 2021, Review of Eudyasmus, with descriptions of a new species from Waigeo Island, Indonesia, and a closely related new genus (Coleoptera: Curculionidae, Molytinae, Eudyasmini), Fragmenta entomologica 53 (2), pp. 377-390 : 379-380

Eudyasmus Pascoe, 1885: 275 .

Type Species. Eudyasmus albertisii Pascoe, 1885 , by monotypy.

Diagnosis. Eudyasmus is similar in appearance to the genus Panopides and the species of Asytesta belonging to the A. compressipes group ( Setliff 2009), but is easily distinguishable by having a flatter and more regular elytral and pronotal profile, by the small median horn-like protrusion at the base of the rostrum (males only), and by the mesoventral receptacle nearly obsolete. The key to the crowned weevil genera in Setliff (2012: 23) will separate Eudyasmus from the other related genera in the crowned weevil group (all of which have a well-developed mesoventral receptacle) except for the new genus described below. See Table 1 for diagnostic features used to distinguish these two closely related genera.

Redescription. Body elongate oval; TL 7.0– 11.3 mm, EW 2.8–5.3 mm; males slightly larger than females on average. Pronotum and elytra subequal in width in dorsal view; humeri not produced. Integument dark reddish-brown to black with extremities lighter in color. Vestiture composed entirely of non-iridescent scales, including small dark brown to black background scales interspersed with evenly distributed longer, suberect, yellowish-brown to white scales that are broadly lanceolate, capitate, or spatulate. Hair-like scales sometimes present on prorostrum, venter, and legs. Pronotum and elytra usually variously marked with white vittae or maculae. Head visible in dorsal view; eyes large, finely faceted, not protruding beyond curvature of head, sub-rounded with posteroventral margin linear, interocular distance narrower than subbasal width of rostrum. Male rostrum compressed, shorter than pronotum, weakly arcuate, more abruptly bent at apex near antennal insertions; female rostrum more evenly curved. Male metarostrum carinate, prorostrum rugose-punctate, bearing small, medial, horn-like protrusion near base; metarostrum smooth in females, finely punctate, prorostrum lacking carinae or basal horn-like protrusion. Antennae inserted beyond middle of rostrum in males, near middle in females. Scape as long as or slightly shorter than funicle in both sexes. Pronotum bell-shaped in dorsal view; disk densely punctate; punctures crowded, evenly distributed, uniform in size; interspaces smooth and glabrous or weakly rugose. Pronotal disk with a very short, thin, glabrous, medial longitudinal carinae at apical margin reaching apical fifth, obliterated well before reaching middle of disk. Postocular lobes well developed with row of vibrissae. Prosternal canal with sides weakly carinate; mesoventral receptacle cup-shaped; open posteriorly, usually covered in long, hair-like scales ( Fig.13a ). Pro- and mesocoxal cavities separated by less than half the width of a coxa, meso- and metacoxal cavities separated by at least half the length of a coxa or more. Metaventrite short- er than abdominal ventrite 1. Scutellum small, oval, wider than long, usually bearing at least some scales. Elytra with sides convex in dorsal view, evenly tapering to apex; with ten visible striae, tenth greatly reduced after basal third, appearing to join the ninth but is traceable to apex. Elytra with small flattened, glabrous area immediately posterior to scutellum on first two intervals of each elytron; elytral interval three weakly granulate and slightly elevated above adjacent intervals; intervals 5–8 not produced or forming a callus at declivity. Metathoracic wings fully developed (as in Fig. 6 ). Sclerolepidia not observed and most likely absent. Legs very long, hind femur capable of exceeding elytral apex by more than half its length, mesofemur also capable of exceeding elytral apex. Profemora edentate, meso- and metafemora with small subapical denticle on venter. Tibia strongly compressed throughout entire length; dorsal margin straight to convex. Protibia straight in apical third, weakly bent ventrad at extreme apex; uncus and premucro well developed; subapex with oblique row of short stout bristles ascending from near uncus to perpendicular angle of apex, forming tooth-like supra-uncal projection (see Fig. 12a, b ). Tarsomere 1 elongate, 1.0–1.2 times longer than tarsomeres 2 and 3 combined; flattened; tarsomere 2 trapezoidal, flattened; tarsomere 3 deeply bilobed; claws simple, free. Ventrite 1 greatly distended, at a much lower plane in lateral view than ventrites 2–5, about as long as remaining ventrites combined, intercoxal process broad; ventrites 2–4 with at least some scales (although they may have glabrous patches in the middle). Male and female genitalia as in Figs. 4–5 .

Distribution. The genus is endemic to the Papuan subregion and is best known from lowland areas of New Guinea and its closest neighboring islands ( Fig. 1 ).

Figures

Fig. 13 – Detail of pro- and mesothorax, ventral view (PSC = prosternal canal, MVR = mesoventral receptacle) – a, Eudyasmus albertisii Pascoe, male; b, Protrachyasmus planidorsis (Heller, 1908) comb. nov., male.

Fig. 6 – Eudyasmus basalis sp. nov., Metathoracic wing.

Fig. 12 – Prothoracic leg details; a, Eudyasmus albertisii Pascoe, 1885, female (PM = premucro, U = uncus, SUP = supra-uncal projection); b, same, male; c, Protrachyasmus planidorsis (Heller, 1908) comb. nov., male.

Fig. 4 – Eudyasmus basalis sp. nov. Male genitalia; a, Penis, lateral view; b, Penis, ventral view; c, Penis, apex in dorsal view; d, Spiculum gastrale; e, Tegmen; f, Tergite VII.

Fig. 5 – Eudyasmus basalis sp. nov., Female genitalia; a, sternite VIII; b, spermatheca, lateral view; c, hemisternites; d, tergite VII; e, tergite VIII.

Fig. 1 – Distribution of the species of Eudyasmus Pascoe, 1885 and Protrachyasmus Setliff gen. nov. Filled in symbols = type locality. Base map from MapCreator 3.0.