Hagnagora richardi Brehm

Brehm, Gunnar, 2015, Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus, ZooKeys 537, pp. 131-156 : 138-139

Taxon classification Animalia Lepidoptera Geometridae

Hagnagora richardi Brehm sp. n. Figs 9, 13

Type material.

Holotype: male (Fig. 9): Ecuador, Loja province, Parque Nacional Podocarpus, Cajanuma, 04°06.85'S, 79°10.47'W, 2916 m, 20 November 2008, G. Brehm leg. (ID 18080, genitalia preparation, barcode sequence 658 bp) (PMJ).

Paratypes: (deposited in CISEC, PMJ, RCGB, ZSM) 4 males, 2 females. 1 female: same as holotype but 04°06.86'S, 79°10.46'W, 2897 m, F. Bodner leg. (ID 15855, barcode sequence 658 bp); 1 female (Fig. 13): Ecuador, Zamora Chinchipe, Reserva Biológica San Francisco, 03°58.72'S, 79°04.44'W, 2180 m, 16 November 2008, F. Bodner leg. (ID 16285, barcode sequence 658 bp); 1 male same as previous but 28 October 1999, D. Süßenbach leg. (BC ZSM Lep 04774, barcode sequence 529 bp); 1 male: same as previous but 03°59.65'S, 79°04.10'W, 2670 m, G. Brehm leg. (ID 17328, barcode sequence 658 bp); 1 male as previous but 03°59.68'S, 79°04.10'W, 2677 m, 18 November 2008 (ID 16119, barcode sequence 658 bp); 1 male as previous but 25 November 2008 (ID 17863, barcode sequence 621 bp).

Description.

As illustrated in Figs 9, 13. The wing length of the holotype (male) is 19 mm. The wing length of a female paratype (Fig. 13) is 21 mm.

Distribution.

Only known from a small region around Podocarpus National Park, provinces Zamora-Chinchipe and Loja, Ecuador, with an observed elevational range of 2180-3021 m a.s.l. Apart from the

Type locality

and nearby sites, specimens were collected at elevations at ca. 3000 m at Cerro Toledo in the same National Park (04°23'S, 79°07'W). However, this record is not fully reliable because genitalia preparation or barcoding was not conducted for these specimens.

Diagnosis.

Closely resembles other species of the Hagnagora anicata clade. On average significantly larger than Hagnagora anicata , but the female has about the same size as Hagnagora hedwigae . The uncus of the male is larger and broader than in Hagnagora anicata . The signum of the bursa copulatrix is less complex than in Hagnagora anicata and Hagnagora hedwigae . Easily distinguishable from Hagnagora marionae by the cream-white colour of the blotches on the forewing. COI barcode: The minimum observed distance to the presumably most closely related species ( Hagnagora marionae ) is 6.6%.

Etymology.

Hagnagora richardi is named in honour of Richard Philipp from Jena, Germany, in recognition of his and his parents’ support for the taxonomy of Neotropical geometrid moths.