Leiurus abdullahbayrami Yağmur, Koç et Kunt, sp., 2009

Yağmur, Ersen Aydın, Koç, Halil & Kunt, Kadir Boğaç, 2009, Description of a new species of Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) from Southеastеrn Turkey, Euscorpius 85 (85), pp. 1-20 : 2-8

Leiurus abdullahbayrami Yağmur, Koç et Kunt, sp.

nov. Figures 1–22 ; Tables 1–2

Records cited in the literature as Leiurus quinquestriatus : TURKEY: Adıyaman Province (Tulga,

1960): National Park of Nemrut Mountain (Crucitti & Cicuzza, 2000). Hatay Province (Kinzelbach, 1984). Kilis Province (Kinzelbach, 1984), Elbeyli District (1,5 km S); Musabeyli District, Aşağı Kalecik Village; Musabeyli District, 1 km W of Hasancalı Village; Musabeyli District, Hasancalı- Yedigöz Villages fork in road; Central District, 1 km E of Küplüce Village; Musabeyli District, 1 km E of Yuvabaşı Village (Yağmur et al., 2007). Mardin Province: Deyrzafran Monastery (Crucitti & Vignoli, 2002) (see Map at Fig. 1 , No. 1). Gaziantep Province: Nizip District, 1 km S of Tanır Village; Nizip District, 2 km NE of Kıratlı Village; Şahinbey District, 1 km south of Kızılhisar Village; Nizip District, 1 km S of Türkyurdu Village; Oğuzeli District, 2 km E of Keçikuyusu Village; Oğuzeli District, 1 km S of Kayacık Village fork in road; Şahinbey District, Sarısalkım Village; Şahinbey District, 2 km N of Sarıkaya Village; Şehitkamil District, 25 th km from Gaziantep to Nizip; Şehitkamil District, Sinan Village, fork in road, 22 th km from Gaziantep to Nizip; Şehitkamil District, Köksalan Village; İslahiye District, 1 km E of Yesemek Village (Yağmur, 2005).

Comparative Material: Leiurus quinquestriatus , 1

♂, 2 ♀♀, EGYPT (without exact locality data) ; 1 ♀, 1

juv., ISRAEL, southern part of Negev Desert , Vadi Hazaz near Sede Boqer (Haluqim Ridge), November 2004, J. Král leg.; 1 ♂ , SYRIA, Bosra , May 1994, D. Modrý leg.

Type Material: Holotype: male, TURKEY, Gaziantep Province: Şahinbey District, a hill 1 km E of Sarısalkım Village, 14 August 2004, 37°05′46.5″N, 37°16′51.3″E, 1029 m asl., E. A. Yağmur leg., steppe area, under rock. Deposited in the Museum of Turkish Arachnology Society , Ankara, MTAS / But:0908-01. (see Map at Fig. 1 , No. 2)

Paratypes (58 females, 36 males, and 50 juveniles): Adıyaman Province : 1 juv., Besni District , 1 km S of Çakırhöyük Village fork in road, 07 July 2006, 37°33′09″N, 37°45′12″ E, 724 asl, E. A. Yağmur & S. Anlaş leg., MTAS / But:0908-02 (3) ; 2 ♀♀, 2 ♂♂, 3 juv., Central District , 1 km W of Aydınoluk Village fork in road, 11 August 2006, 37°46′08″E, 38°20′54″N, E. A. Yağmur & A. Avcı leg., MTAS / But:0908-07 (4) ; 3 ♀♀, 1 ♂, 5 juv., Central District, Gökçay Village, Konakdere Hamlet , 29 July 2007, 37°57′24.9″N, 38°17′55.3″E, 1155 asl, E. A. Yağmur & A. Avcı leg., MTAS / But:0908-22 (5) ; 1 juv., Kahta District, National Park of Nemrut Mountain , 1 km N of the gate, 09 June 2007, 37°56′49″N, 38°44′47″E, 1535 m asl, E. A. Yağmur, G. Çalışır leg., MTAS / But:0908-13 (6) ; 1 ♀, Tut District, 2 km SW of Meryemuşağı Village fork in road, Hacıvakı Suyu area , 03 June 2007, 37°48′31″N, 37°51′49″E, 1143 asl, E. A. Yağmur & M. Özkörük leg., MTAS / But:0908- 23 (7) ; 4 ♂♂, Tut District , 4 km E of Kaşlıca Village, 08 June 2007, 37°48′34″N, 37°59′21″E, 1183 m asl, E. A. Yağmur & G. Çalışır leg., MTAS / But:0908-12 (8). Gaziantep Province : 2 ♂♂, İslahiye District , a hill E of Yesmek Village, 14 May 2005, 36°54′00″N, 36° 44′43″E, 487 m asl, E. A. Yağmur & M. Pehlivanoğlu leg., MTAS / But:0908-68 (9) ; 1 ♂, Nizip District , 1 km S of Tanır Village, 30 May 2003, 37°06′34″N, 37°49′57″E, 529 m asl, E. A. Yağmur & S. Kesmezoğlu leg., MTAS / But:0908-45 (10) ; 1 ♀, Nizip District , 10 km W of Türkyurdu Village, 15 September 2007, 37°02′44″N, 37°34′23″E, 804 m asl, E. A. Yağmur, E. Ulupınar leg., FKCP (09F61-2) (11) ; 1 juv., Nizip District, Türkyurdu Village , 20 January 2008, 37° 00′33″N, 37°37′42″E, 703 asl, E.A. Yağmur & M. Yalçın leg., MTAS / But:0908-36 (12) ; 1 juv., Oğuzeli District, Çaybeyi Village , 23 May 2008, 36°47′48″N, 37°35′15″E, M. Z. Yıldız leg., MTAS / But:0908-37 (13) ; 1 ♀, 1 ♂, 1 juv., Oğuzeli District , 1 km NE of Çaybaşı Village, 25 June 2006, 36°47′47″N, 37°35′15″E, 546 m asl, E. A. Yağmur & M. Özkörük leg., MTAS / But:0908- 44 (14) ; 1 ♀, Oğuzeli District , 1 km S of Kayacık Village fork in road, 11 September 2004, 36°50′41″N, 37°34′29″E, 619 m asl, E. A. Yağmur & M. Yalçın leg., MTAS / But:0908-47 (15) ; 1 ♂, Şahinbey District, 1 km W of Güllüce Village, Lower slop of Ellezi Hill , 29 July 2006, 36°56′38″N, 37°05′12″E, 822 m asl, E.A. Yağmur & M. Yalçın leg., MTAS / But:0908-40 (16) ; 1 ♂, Şahinbey District , 2 km SW of Gerciğin Village, 21 May 2006, 37°02′11″N, 37°15′00″E, 1034 m asl, E.A. Yağmur leg., MTAS / But:0908-41 (17) ; 1 ♂, Şahinbey District , 2 km N of Sarıkaya Village, 07 May 2005, 37°07′06″N, 37°03′07″E, 1178 m asl, E. A. Yağmur, M. Yalçın & F. Değirmenci leg., MTAS / But:0908-46 (18) ; 1 ♀, 2 ♂♂, Şahinbey District, Sarısalkım Village, Şirvan Hill , 06 September 2003, 37°05′27″N, 37°15′52″E, 1076 m asl, E. A. Yağmur leg., MTAS / But:0908-50 (19) ; 1 ♀, Şahinbey District , a hill E of Sarısalkım Village, 30 July 2003, 37°05′47″N, 37°16′51″E, 1024 m asl, E. A. Yağmur leg., MTAS / But:0908-51 (19) ; 1 ♀, 1 ♂, Şahinbey District , a hill E of Sarısalkım Village, 12 April 2004, 37°05′47″N, 37°16′51″E, 1024 m asl, E. A. Yağmur leg., MTAS / But:0908-53 (19) ; 1 ♀, Şahinbey District , a hill E of Sarısalkım Village, 06 September 2004, 37°05′47″N, 37°16′51″E, 1024 m asl, E. A. Yağmur leg., MTAS / But:0908-54 (19) ; 1 ♀, 4 ♂♂, Şahinbey District , a hill E of Sarısalkım Village, 14 August 2004, 37°05′47″N, 37°16′51″E, 1024 m asl, E. A. Yağmur leg., MTAS / But:0908-59 (19) ; 1 ♂, Şahinbey District , 2 km S of Sarısalkım Village, brick factory, 10 July 2003, 37°04′57″N, 37°17′11″E, 943 m asl, E. A. Yağmur leg., MTAS / But:0908-60 (19) ; 1 ♀, 1 ♂, Şahinbey District, Sarısalkım Village , 0,5 km N of school, 12 April 2004, 37°06′16″N, 37°16′48″E, 1034 m asl, E. A. Yağmur leg., MTAS / But:0908-62 (19) ; 1 ♀, 1 ♂, Şahinbey District , 1 km S of Kızılhisar Village, 14 September 2004, 36°59′52″N, 37°18′17″E, 900 m asl, E. A. Yağmur & C. Toprak leg., MTAS / But:0908-65 (20) ; 1 ♂, Şehitkamil District , 1 km S of Eski Şarkaya Village, 23 June 2007, 37°12′44.4″N, 37°07′45.5″E, 1000 m asl, E.A. Yağmur & M. Yalçın leg., FKCP (09F96-97) ; 2 ♂♂, 2 juv., Şehitkamil District , 1 km S of Eski Şarkaya Village, 23 June 2007, 37°12′44.4″N, 37°07′45.5″E, 1000 m asl, E.A. Yağmur & M. Yalçın leg., MTAS / But:0908-27 (21) ; 1 ♀, 1 ♂, 4 juv., Şehitkamil District , 2 km E of Yamaçoba Village, 01 July 2007, 37°10′27.3″N, 37°05′33.1″E, 1099 m asl, E.A. Yağmur & M. Yalçın leg., MTAS / But:0908-33 (22) ; 1 ♀, Şehitkamil District, İncesu Village , 04 May 2008, 37°13′16″N, 37°18′05″E, 942 m asl, E. A. Yağmur & E. Tezcan leg., MTAS / But:0908-34 (23) ; 1 juv., Şehitkamil District, Tuğlu Village , 11 May 2007, 37°10′56″N, 37°31′51″E, 777 m asl, E.A. Yağmur & M. Yalçın leg., MTAS / But:0908-35 (24) ; 1 ♀, 1 juv., Şehitkamil District , 1,5 km NW of Suboğazı Village, 11 May 2007, 37°09′35″N, 37°28′51″E, 814 m asl, H. Koç leg., MTAS / But:0908-39 (25) ; 1 ♀, Şehitkamil District, Sinan Village fork in road, 22 th km from Gaziantep to Nizip , 14 September 2004, 37°02′31″N, 37°35′48″E, 746 m asl, E. A. Yağmur & M. Özkörük leg., MTAS / But:0908-63 (26) ; 1 ♀, Şehitkamil District, Köksalan Village , 09 April 2005, 37°15′44″N, 37° 14′32″E, 956 m asl, E. A. Yağmur, M. Yalçın & F. Değirmenci leg., MTAS / But:0908-66 (27). Hatay Province : 2 juv., Reyhanlı District, Oğulpınar Village , 28 May 2007, 36°15′13″N, 36°40′12″E, 316 m asl, H. Koç, A. V. Gromov leg., MTAS / But:0908-70 (27) ; 2 ♀♀, 1 ♂, 3 juv., Reyhanlı District, Kavalcık Village . 26 May 2007, 36°14′45.6″N, 39°37′16.1″E, H. Koç, A. V. Gromov leg., MTAS / But:0908-76 (28) ; 3 ♀♀, Reyhanlı District, Kavalcık Village . 27 May 2007, 36°14′45.6″N, 39°37′16.1″E, H. Koç, A. V. Gromov leg., MTAS / But:0908-79 (28) ; 2 ♀♀, 4 juv., Reyhanlı District, Bükülmez Village , 27 May 2007, 36°18′15″N, 36° 38′12″E, 157 m asl, H. Koç leg., MTAS / But:0908-85 (29) ; 4 ♀♀, 1 ♂, 2 juv., Reyhanlı District , 1 km W of Çakıryiğit Village, 14 April 2007, 36°15′42″N, 36°36′22″E, 225 m asl, E. A. Yağmur, M. Yalçın, G. Çalışır leg., MTAS / But:0908-92 (30) ; 3 juv., Reyhanlı District , 1 km E of Fevzi Paşa Village, 14 April 2007, 36°15′52″N, 36°38′27″E, 256 m asl, E. A. Yağmur, M. Yalçın, G. Çalışır leg., MTAS / But:0908-95 (31). Kahramanmaraş Province : 1 ♀, Pazarcık District , 0.5 km N of Hörük Village fork in road, 24 June 2007, 37°30′10″N, 37°33′46″E, 795 m asl, E. A. Yağmur & M. Yalçın leg., MTAS / But:0908-96 (32) ; 2 juv., Türkoğlu District, Evri Village , 23 June 2007, 37°16′13″N, 37°08′33″E, 720 m asl, E.A. Yağmur & M. Yalçın leg., MTAS / But:0908-98 (33). Kilis Province : 1 ♀, 1 ♂, Central District, 2 km E of Küplüce Village, 23 April 2007, 36°44′92″N, 37°15′19″E, 624 m asl, E. A. Yağmur & M. Z. Yıldız leg., Zoology Collection of Ege University Museum, İzmir, Turkey ( ZDEU); 1 ♀, 1 juv., Central District , 2 km E of Küplüce Village, 23 April 2007, 36°44′92″N, 37°15′19″E, 624 m asl, E. A. Yağmur & M. Z. Yıldız leg., OUMNH-2009-022 , OUMNH-2009-023 ; 1 ♀, 2 juv., Central District , 2 km E of Küplüce Village, 23 April 2007, 36°44′92″N, 37°15′19″E, 624 m asl, E. A. Yağmur & M. Z. Yıldız leg., FKCP (09F61a) ; 4 ♀♀, 1 ♂, 6 juv., Central District , 2 km E of Küplüce Village, 23 April 2007, 36°44′92″N, 37°15′19″E, 624 m asl, E. A. Yağmur & M. Z. Yıldız leg., MTAS /But:0908-110 (35); 4 ♀♀, 1 juv., Central District, 1 km E of Küplüce Village, 29 April 2006, 36°44′92″N, 37°15′19″E, 624 m asl, E. A. Yağmur & M. Yalçın leg., MTAS / But:0908- 115 (35) ; 1 ♀, Central District, Çörten Village , 27 June 2007, 36°46′35″N, 37°18′00″E, M. Z. Yıldız leg., MTAS / But:0908-123 (36) ; 1 ♀, 1,5 km S of Elbeyli District, Çamurluhöyük mound (tumulus), 30 April 2006, 36°39′36″N, 37°28′31″E, 525 m asl, E. A. Yağmur & M. Özkörük leg., MTAS / But:0908-125 (37) ; 1 ♀, Musabeyli District , 3 km SW of Hasancalı Village, 27 April 2006, 36°52′47.2″N, 36°47′42.4″E, 755 m asl, E. A. Yağmur & M. Özkörük leg., MTAS / But:0908-116 (38) ; 1 ♀, 2 ♂♂, 2 juv., Musabeyli District , 3 km SW of Hasancalı Village, 26 May 2007, 36°52′47″N, 36°47′42″E, 755 asl, H. Koç & A. V. Gromov leg., MTAS / But:0908-121 (38) ; 2 ♀♀, 1 ♂, Musabeyli District , 3 km SW of Hasancalı Village, 30 May 2007, 36°52′47″N, 36°47′42″E, 755 asl, H. Koç & A. V. Gromov leg., MTAS / But:0908-122 (38) ; 1 ♀, Polateli District , 2 km N of Polateli fork in road, 01 March 2008, 36°46′06,0″N, 37°04′17,1″E, E. A. Yağmur & E. Ulupınar leg., MTAS / But:0908-99 (39) ; 1 ♀, Polateli District, Aşağı Kalecik Village , 22 April 2006, E. A. Yağmur, M. Yalçın & M. Özkörük leg., MTAS / But:0908-124 (40) ; 1 ♀, Musabeyli District, Hasancalı Village-Yedigöz Village fork in road, 27 April 2006, 36°51′22″N, 37°49′46″E, 663 m asl, E. A. Yağmur & M. Özkörük leg., MTAS / But:0908-126 (41). Şanlıurfa Province : 1 ♀, 1 ♂, Birecik District , 2 km N of Yukarı Habib Village, 24 April 2009, 37°08′49″N, 37°59′56″E, 670 m asl, E. A. Yağmur & V. Ülgezer leg., MTAS / But:0908-128 (42) ; 1 juv., Halfeti District , 2 km S of Yeşilözen Village, 24 April 2009, 37°11′28.9″N, 37°58′08.8″E, E. A. Yağmur & V. Ülgezer leg., MTAS / But:0908-133 (43) ; 3 ♀♀, Harran District , 2 km NE of Şuayipşehir Village, 06 May 2006, 36°52′36.9″N, 39°22′18.9″E, 506 m asl, E. A. Yağmur & M. Z. Yıldız leg., MTAS / But:0908-131 (44) ; 1 ♀, Harran District , 2 km N of Şuayipşehir Village, 25 April 2009, 36°52′36.9″N, 39°22′18.9″E, 506 m asl, E. A. Yağmur & V. Ülgezer leg., MTAS / But:0908-132 (44) .

Derivatio nominis: The new species is named after Prof. Dr. Abdullah Bayram (University of Kırıkkale, Turkey), a specialist on Araneae, who has conducted important arachnological research in Turkey.

Diagnosis: Leiurus abdullahbayrami sp. nov. ( Figs. 2, 3 and 4 ) belongs to the genus Leiurus Ehrenberg : its tergites I and II bear five carinae and centrolateral and fused posteriomedian carinae form a lyre-shaped figure ( Figs. 5–6 ). Because this species was previously incorrectly identifed as Leiurus quinquestriatus (Ehrenberg, 1828) , it is compared with other species of Leiurus and particularly with L. quinquestriatus . It can be distinguished from L. quinquestriatus by the following characters:

(a) background color of prosoma, mesosoma and segment V of metasoma of the new species are black, appendages are yellowish; note that Leiurus jordanensis also has a generally blackish coloration except for its vesicle;

(b) fixed finger and movable fingers of pedipalps have 11–12 (generally 11, rarely 12) oblique denticle rows ( Fig. 16 ), whereas L. quinquestriatus has 12–13 (generally 12, rarely 13) rows. Movable fingers of pedipalps have 13 rows in L. savanicola , 12 rows in L. jordanensis and 11 – 12 rows in L. nasheri .

(c) ventrolateral carinae of metasomal segment V are armed with large and rounded granules ( Figs. 7 and 14 ), whereas the same carinae in L. quinquestriatus bear smaller and rounded lobes. The size of ventrolateral carinal denticles of metasoma V is variable in L. quinquestriatus . We have examined a large number of specimens but the new species does not exhibit considerable variation in ventrolateral carinae of the fifth segment. Hendrixson (2006) illustrated variation in lobe size in ventrolateral carinae of metasomal segment V. All our specimens bear large and round lobes in ventrolateral carinae of metasomal segment V compared with all populations illustrated in Hendrixson (2006) ( Fig. 7 ). The ventrolateral carinae of the metasomal segment V in Leiurus jordanensis Lourenço et al., 2002 are composed of spinoid lobes.

(d) the anal arch on the metasomal segment V in the new species comprises 2–3 rounded lobes laterally and 5–6 small lobes posteriorly ( Figs. 7, 8, 9 , and 15), whereas the anal arch of L. quinquestriatus is composed of 3 partly spinoid lobes laterally and 9–10 smaller lobes posteriorly. Leiurus can dig in hard soils using their legs and metasoma V, and circum-anal dentition and ventrolateral dentition can be worn out or rounded. For this reason, we have examined also juvenile specimens. We still observed large and rounded lobes on ventrolateral carinae of metasoma V and rounded lobes on lateral sides of anal arch. Therefore, these structures were treated as characteristic features for Leiurus abdullahbayrami sp. nov.

(e) in the new species, trichobothrium db on the fixed finger of the pedipalp is located between trichobothria est and esb ( Fig. 17 ). This trichobothria pattern resembles Leiurus nasheri Kovařík, 2007 ; however, trichobothrium db on the fixed finger of the pedipalp is located between trichobothria et and est in L. quinquestiatus , L. jordanensis and L. savanicola .

(f) the pedipalp of L. quinquestriatus is more slender than in Leiurus abdullahbayrami sp. nov. In particular, movable finger of Leiurus abdullahbayrami sp. nov. is shorter than in L. quinquestriatus ( Fig. 11 ). The pedipalp average length/width ratio is 4. 72 in the males and 4.64 in the females of Leiurus abdullahbayrami sp. nov. When this ratio is calculated for L. quinquestriatus from published tables according to Levy & Amitai (1980), it is 5. 42 in males, 5. 64 in females; according to Lourenço et al. (2002), it is 6.11 in females; according to Lourenço et al. (2006), it is 7.76 in males. Among the specimens investigated, the ratio in L. q. quinquestriatus for male was 7.08, and for a female, 6.26; in L. q. hebraeus, 5.00 for a male, 5.74 for a female. Additionally, according to tables in Lourenço et al. (2002), in females of L. jordanensis the ratio is 8.47; in males of L. savanicola it is 6.95 (Lourenço et al., 2006); in females of L. nasheri , it is 4.82 (Kovařík, 2007). According to the literature, the length/width ratio of the pedipalp of Leiurus abdullahbayrami sp. nov. is significantly different from L. quinquestriatus , L. jordanensis and L. savanicola ; on the other hand, it is close to L. nasheri .

(g) sternites IV–VI of Leiurus abdullahbayrami sp. nov. are smooth, but they are granulated in L. quinquestriatus .

(h) the sting of L. quinquestiatus is slightly more curved than sting of Leiurus abdullahbayrami sp. nov. ( Fig. 10 )

The new species has pairs of short spines on the ventral side of tarsus of the legs I–IV ( Figs. 20 and 21 ), similarly to L. quinquestriatus , L. jordanensis and L. savanicola . However, L. nasheri bears long paired ventral setae on the first and second tarsus, quite different from other Leiurus species.

Description of male holotype (for morphometric measurements see Table 1.)

Coloration: Body black or blackish brown, appendages light yellow ( Figs. 3 and 4 ). Prosoma:

carapace blackish-brown with semicircular yellow zone anteriorly ( Figs. 3 and 4 ), or this area yellowish black ( Fig. 6 ) and a pair of yellowish black spots laterally and posteriorly ( Figs. 3 , 4 , and 5). Lateral eyes surrounded by black pigmentation. Anterior median carinae completely black. Mesosoma: black or blackish-brown. Metasoma: segments I to IV yellowish; ventral submedian carinae distinctly black on segment IV, slightly black on segments II and III; segment V is black, sometimes lighter in adults, always of darker coloration in subadults. Vesicle pale yellow; aculeus yellowish at base and reddish-brown at its extremity. Chelicerae yellowish with little dark reticulated spots; denticles reddish-brown. Pedipalps: dark brown overall; rows of denticles on dentate margins of fingers reddish-brown. Leg femur and patella dark yellow; other segments light yellow.

Figures

Figure 1: Sampling localities of Leiurus abdullahbayrami sp. nov. in Eastern Mediterranean Region and Southeastern Anatolia Region, Turkey.

Figures 2–3: Leiurus abdullahbayrami sp. nov. 2 (top). In a natural habitat (Yukarı Habip Village, Birecik District, Şanlıurfa Province). 3 (bottom). In captivity (a specimen from Gaziantep Province).

Figure 4: Leiurus abdullahbayrami sp. nov. male holotype (left) and a female paratype (Gökçay Village, Adıyaman Province) (right).

Figures 5–6: Leiurus abdullahbayrami sp. nov. carapace. 5. Female paratype (Gökçay Village, Adıyaman Province). 6. Male paratype (2 km E of Yamaçoba Village, Şehitkamil District, Gaziantep Province).

Figure 7–9: Leiurus abdullahbayrami sp. nov. 7. Ventrolateral carinae of metasomal segment V. 8. Anal arch, laterally. 9. Anal arch, ventrally.

Figure 10: Stings of female Leiurus quinquestriatus (left) (Israel) and female L. abdullahbayrami sp. nov. (right) (a paratype from Adıyaman Province).

Figure 11: Pedipalps of female Leiurus quinquestriatus (left) (Israel) and female Leiurus abdullahbayrami sp. nov. (right) (a paratype from Adıyaman Province).

Figure 12–13: Leiurus abdullahbayrami sp. nov., coxosternal area. 12. Female paratype (Adıyaman Province). 13. Male holotype.

Figure 14–19: Leiurus abdullahbayrami sp. nov. 14. Lobes of the ventrolateral carinae of metasomal segment V. 15. Anal arch, ventrally. 16. Dentition of movable finger. 17. Trichobothria pattern of chela, external view only. 18. Trichobothria pattern of patella, dorsal view only. 19. Trichobothria pattern of femur, dorsal view only.

Figure 20–21: Leiurus abdullahbayrami sp. nov. 20. Tarsus of leg III. 21. Tarsus of leg IV.

Figure 22: Leiurus abdullahbayrami sp. nov. color variation among three populations: Hatay Province (left), Şanlıurfa Province (center); Gaziantep, Adıyaman, and Kilis Provinces (right).