Clivina komareki KULT , 1951

Balkenohl, Michael, 2021, The Clivina komareki-species group from Asia with the description of a new species from the Philippines (Coleoptera, Carabidae, Scaritinae), Linzer biologische Beiträge 53 (2), pp. 535-543 : 537-539

Clivina komareki KULT, 1951 ( Figs 1 , 3)

T y p e m a t e r i a l: Holotype: ♁, with labels and data:white, printed in black, "Simbang Huon Golf" / "N Guinea Biro 1898" / white, handwritten in pencil "XIX" / red, handwritten in black ink "coll. m." and with red pencil "2" / white, handwritten with pencil " C. Komareki " / red, printed in black " HOLOTYPUS? Coll. K. KULT Dostal fecit 1999" / white, handwritten in black ink and printed " Clivina komareki KULT 1951 det. Dr. A. Dostal 1999" (CDW).

N o t e: The species was characterized with reasonable accuracy in an one-page description ( KULT 1951, p. 31, 32). Based on one other specimen, DARLINGTON (1962) provided additional information and a line sketch of the anterior margin of the clypeus. The holotype studied is in good condition but with two terminal tarsomeres of the left mesotibia missing.

The specimen does not carry a label with the direct writing of the word "typus", but a red "coll. m." label. In 1999 and after the acquisition of the collection Kult, Dostal added a red holotype label to the specimen indicated with "fecit". On the label he added a question mark, obviously to indicate some reluctance.

After investigating the specimen and comparing it with the description in detail, I am convinced it is the holotype Kult based the description on due to the following reasons: The holotype fits exactly with the description and as well with the key provided by Kult (1951). The locality the specimen was found is correctly indicated. The species is conspicuously different from the other Oriental species. In addition, it is well documented that the collector Lajos Biró collected at the type locality in New Guinea in 1898 ( HORN et al. 1990).

However, dissection revealed the holotype is a male and not a female as indicated in the description by KULT (1951, p. 32).

Because the description does not include figures and some characters are not described in detail, a full redescripten is given.

D i a g n o s i s: See recognition of the Clivina komareki- species group.

R e d e s c r i p t i o n:

Measurements: Holotype: Body length 5.57 mm, width 1.63 mm; ratio length/width of pronotum 0.96; ratio length/width of elytra 1.85.

Colour: Glossy. Piceous. Legs uniformly fuscous, antenna and mandible slightly paler fuscous.

Head: Nearly a third narrower than pronotum. Clypeus straight anteriorly; wing wide, prominent, distinctly more protruding anteriorly than clypeus, fused with clypeus; supraantennal plate wide, convex in dorsal and frontal view; clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level as obtuse carina, separated from wing by a notch. Clypeus transverse, moderately convex, separated from frons by a wide and moderately deep furrow with irregular punctures. Frons moderately convex, with triangle-like depression at middle. Clypeus, frons, and supraantennal plates smooth, separated by deep moderately wide furrow continuing posteriorly up to posterior eye-level as wide supraorbital furrow. Two supraorbital setigerous punctures situated at mid and posterior eye level. Neck constriction developed as distinct step, with uninterrupted row of punctures of moderate size. Eye hemispherical, projecting laterally. Gena small. Antenna reaching up to posterior seta of pronotum, antennomeres four to ten of moderate length (ratio L/W around 1.3). Labrum straight, indistinctly reticulated, six setose. Mandible wide. Mentum with lateral lobe like an askew trapezium, margin straight anteriorly, median tooth short, keeled.

Pronotum: Disk slightly convex in lateral view, moderately convex in frontal view. Quadrate, slightly wider than long. Anterior margin indistinctly emarginated. Reflexed lateral margin smooth, straight at middle, slightly attenuating in anterior half, widest in posterior third; anterior angle obtuse but distinct; posterior angle marked by distinct tooth, laterally projecting as far as lateral margin in posterior third. Lateral channel indistinctly foveolate, of equal width in its whole length. Median line narrow, line-like, joining anterior transverse line and basal constriction; anterior transverse line deeper than median line, not reaching lateral channel. Surface with microscopic punctures, with group of few moderately sized punctures laterally in basal third, with few transverse wrinkles laterally and at base.

Elytron: Disk slightly convex in lateral view, distinctly convex on frontal view. Elytra concavely impressed in apical quarter (best visible in anterior-frontal view at an angle of 45°). Outline elongate, nearly twice as long as wide, long oval, with maximum width at middle. Humerus rounded but distinct, without humeral tooth. Reflexed lateral margin nearly smooth. Lateral channel wide. Scutellar striole indistinct; with distinct setigerous puncture at base of first stria. Striae deep, indistinctly punctuate, one to four free at base, five to seven joining at humerus, one running up to apex, two ending free at apex, three and four, and five and six joining apically. Intervals distinctly convex, third to fifth more elevated at base, seventh wider and more elevated apically. Third interval with three small setigerous punctures. Surface of intervals glossy, striae indistinctly reticulated.

Hind wing: Fully developed.

Lower surface: Proepisternum covered with slight punctures. All terminal sternites of abdomen laterally with moderately sized punctures. The two setigerous punctures on each side of the abdominal sternum VII are very widely separated.

Legs: Legs covered with longitudinal reticulation. Protibia with three spines of moderate length, not sulcate dorsally, movable spur regularly arcuate. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with four fine seta at the inner side.

Male genitalia (Fig. 3): Median lobe regularly slender, moderately arcuate in basal half, apex developed as moderately slender arcuate shank, distinctly arcuate at tip pf apex. Endophallus with central group of spines, spines slender. Parameres slender, slightly distorted, acuminated in apical half, apex pointed, at apex with one moderately sized seta and one minute seta preapically.

Female genitalia: Unknown.

Variation: Unknown.

D i s t r i b u t i o n: Known from the type locality Simbang at the Huon Golf in Papua New Guinea and from Hollandia, New Guinea.

Figures

Figs 1-2: Clivina species, habitus,dorsal view; (1)C. komareki KULT, holotype; (2) C. excisa nov.sp., holotype.