Leiobunum rupestre (Herbst, 1790)

Martens, Jochen & Schönhofer, Axel L., 2016, The Leiobunum rupestre species group: resolving the taxonomy of four widespread European taxa (Opiliones: Sclerosomatidae), European Journal of Taxonomy 216 (216), pp. 1-35 : 31-35

Leiobunum rupestre (Herbst, 1790)

AUSTRIA: Vorarlberg: 1 ♁, Übersaxen, Baschghöhle, 785 m, 47.25° N, 9.67° E, leg. October (Breuss 1995); Salzburg (all records if not otherwise indicated refer to Ausobsky 1987), Bergheim bei Salzburg, Hochgitzen, 47.850° N, 13.034° E; Buchberg, 47.942° N, 13.092° E; Haunsberg, 47.92° N, 12.99° E; Heuberg, 47.82° N, 13.11° E; Moosachtal bei St. Georgen an der Salzach, 47.98° N, 12.88° E; Plain, Plainberg, 47.837° N, 13.044°E; Salzburg, Festungsberg, 47.795°N, 13.048° E; Salzburg, Kapuzinerberg, 47.803° N, 13.056° E; Salzburg, Mönchsberg, 47.799° N, 13.039° E; Salzburger Becken, Elsbethen, Elsbethener Hügel, 47.76° N, 13.10° E, Georgenberg near Kuchl, 47.635° N, 13.161° E; Hellbrunner Berg, 47.758° N, 13.066° E; Morzger Hügel, 47.768° N, 13.055° E; Tannberg, 47.973° N, 13.188° E; Tiefstein gorge near Schleedorf, 47.947° N, 13.158° E; Stubachtal, Weissee, 47.16° N, 12.64° E (Mazzucco 1967); 4 ♁♁, 2 ♀♀, Stubachtal, N Enzingerboden, 1420–1460 m, 47.184° N, 12.625° E, leg. 7 Oct. 1966 ( CJM 496); 11 ♁♁, 5 ♀♀, Pinzgau, Kaprun, entrance of Sigmund-Thun-Klamm (Alp12_04), 800 m, 47.258517° N, 12.738571° E, leg. 10 Sep. 2012 ( AXLS 1283); Oberösterreich, Weilhartsforst, 48.16° N, 12.91° E (Ausobsky 1987); 1 ♁, vicinity of Stifterherberge at Bärenstein, 900–950 m, upper coniferous forest, 48.660° N, 13.991° E ( SMF 16343/1); 1 ♁, Steiermark, Koralpe, Soboth, 900 m, wet slopes near stream, Salix and Abies , 46.67650° N, 15.08331° E, leg. 30 Aug. 2009 ( AXLS 277); 1 ♁, Gesäuse, coniferous forest, 47.58° N, 14.64° E, leg. 16 Sep. unknown jear ( ZMB 12640); Kärnten, Baldersdorf, way to Egelsee, 46.78° N, 13.56° E (Komposch 2008); Saualpe, Löllingbach, 1100 m, 46.55° N, 14.37° E (Komposch & Gruber 2004); 5 juv., Flaschberg near Oberdrauburg, 46.75° N, 12.94° E, leg. 28 Jul. 1982 (Kofler & Mildner 1986); Klagenfurt, Sattnitz, boulders betw. Bad and Hanswirt, leg. 3 Sep. 1985 (Kofler & Mildner 1986); Klagenfurt, Sattnitz, Höllgraben, 46.49° N, 14.28° E, leg. 10 Sep. 1985 (Kofler & Mildner 1986); 5 ♁♁, 1 ♀, near Klebas, road 111, subway, 46.696° N, 12.802° E, leg. 1 Sep. 2012 ( AXLS 1356); Ruin Obertrixen, 46.702° N, 14.575° E, leg. 19 Aug. 1985 (Kofler & Mildner 1986); St. Leonhard near Liebenfels, 46.74° N, 14.26° E, leg. 30 Jul. 1985 (Kofler & Mildner 1986); 1 ♁, Tröpolach, Gailtal, 900 m, 46.62° N, 13.28° E, leg. 30 Sep. 1975 ( CJM 1411); Ulrichsberg, 46.71° N, 14.31° E, leg. 3 Oct. 1968 (Kofler & Mildner 1986); 6 ♁♁, 2 ♀♀, N Graz, Hinterreitenwald, SE Glettweber, Glettbach, 870 m, 47.201° N, 15.428° E, leg. 14 Sep. 2012 ( AXLS 1301); Hohe Tauern, Gössnitztal (Gossnitz=Goessnitz), 1700 m, 47.031° N, 12.792° E (Komposch & Gruber 1999); Niederösterreich, Lunz am See, forests arround Untersee, 47.85° N, 15.05° E (Thaler 1963); 1 ♁, Semmering (M2), 47.63° N, 15.83° E, leg. Oct. 1899 ( ZMB 12644); 1 ♁, Tirol, Innsbruck, in the city, Weierburggasse, 730–800 m, at housewall, 47.278° N, 11.396° E, leg. 27 Sep.–1 Oct. 1986 ( CJM 3523); Innsbruck surroundings, Picea forests S of Inn River, Husslhof-Natters-Amras-Igls-Ampass-Hall, 580 m, in moss (Stipperger 1928); 1 ex., Forellenhof near Lavant, at limestone boulders, 46.79° N, 12.84° E, leg. 7 Sep. 1971 (Kofler 1984); 3 juv., Nikolsdorf, Tal-Trasse, right side of Drau River, 46.78° N, 12.91° E, leg. 28 Jul. 1982 (Kofler 1984); 1 ♀, Nörsach (Kärntner Grenze), 46.78° N, 12.93° E, leg. 28 Aug. 1969 (Kofler 1984); Villgrater Tal (GPS not accurate), 46.80° N, 12.38° E (Werner 1931); 1 ♀, Virgental, Iselschlucht near Welzelach, 47.017° N, 12.413° E, leg. 2 Aug. 1978 (Kofler 1984); Windisch-Matrei, 47.00° N, 12.53° E (Dalla-Torre 1882); Bärenbadalm above Achensee, 1400–1500 m, caves in Picea forest, 47.42° N, 11.70° E (Stipperger 1928); Hungerburg, 860 m, Sand- and Schotterquarry at border of Picea forest, 47.29° N, 11.40° E (Stipperger 1928); Maldontal, betw. Boden and Bschlabs, 1350 m, Fichten- und Föhrenwald, 47.30° N, 10.59° E (Stipperger 1928); Stallental, 800–1600 m, Picea forest with Fagus , 47.38° N, 11.69° E (Stipperger 1928); Kaunertal from Kaltenbrunn and Feuchten to Gepatschhaus (coordinates inaccurate), 1260–1700 m, Picea forest, humid cliffs along stream and path, 47.05° N, 10.75° E (Stipperger 1928); Oberbergtal from Milders im Stubai to Stöcklenalm (coordinates inaccurate), 1100–1600 m, Picea forest, at boulders, 47.12° N, 11.32° E (Stipperger 1928); S St. Anton am Arlberg, 1310 m, cliffs in Picea forest, 47.13° N, 10.27° E (Stipperger 1928); Schwaz, betw. Schloss Freundsberg and Rodlhütte, 540–1600 m, Picea forest mixed with Abies and Fagus , under leaf litter, in moss, at stones and truncs, 47.34° N, 11.71° E (Stipperger 1928); Sellraintal betw. Kematen, Gries and Praxmar, 610–1600 m, humid, shaded cliffs and moss at stream in Picea forest, 47.17° N, 11.14° E (Stipperger 1928); between Pettneu and Malfontal, 1200–1300 m, Picea forest, 47.140° N, 10.343° E (Stipperger 1928); betw. Volders and Volderer Wildbad, 600–1100 m, wet Picea forest with Alnus ; 3 ♁♁, 47.27° N, 11.56° E (Stipperger 1928); Lienz (GPS not accurate), 46.83° N, 12.76° E, (SMFRII/1184, Roewer 1957); 3 ♁♁, 1 ♀, Lienzer Dolomiten, Hohenstein (SMFRII/701, Roewer 1957); Burgenland, 1 ♁, southern Leithagebirge, „Kürschnergrube” (abandoned chalk quarry) N of Eisenstadt, 435 m, in man made cavity at steep humid cliff, 47.86° N, 16.52° E, leg. 7 Nov. 1996 ( NMW 17922, Gruber 2000).

BOSNIA AND HERZEGOVINA: 11 ♁♁, 1 ♀, Jaice (=Jajce), tufa walls, waterfall, 44.342° N, 17.266° E ( ZMB 12639); 2 ♁♁, Kupreška vrata, Bugojno, 1384 m, 44.00° N, 17.31° E (Novak 2005); Travnik, 44.22° N, 17.66° E (Martens 1978); 2 ♁♁, Trebević (National Park and mountain S of Sarajevo, coordinates accurate), Corylus forest, 43.8° N, 18.45° E, leg. Sep.–Oct. ( ZMB 12638); 4 ♁♁, 1 juv., Vilinska jama cave, 42.73° N, 17.89° E, leg. 1916 (Novak 2005).

CROATIA: 1 ♁, Bijeli vrh, Vrhovine, 1074 m, 44.84° N, 15.44° E (Novak 2004b); 1 juv., Kamengrad, Papuk Mt., beside the Orljava river, 528m, 45.53° N, 17.50° E (Novak 2004b); 5 ♁♁, 2 ♀♀, 2 juv., Kamenica Skradnička, beside the Rudnica brook ( WL 30), 260 m, 45.20° N, 15.45° E, leg. May 1985 (Novak 2004b); 3 ♁♁, 1 ♀, Kameniti vrh Mt., Plitvice, 934 m, 44.88° N, 15.57° E (Novak 2004b); 1 ♁, Klanjec, 170 m, 46.05° N, 15.74° E (Babić 1916); 1 ♁, Krapina, 290 m, 46.16° N, 15.87° E, leg. Aug. 1914 (Babić 1916); Križevci, 140 m, 46.03° N, 16.52° E (Daday 1896), orig. det. as Liobunum glabrum ); 2 ♁♁, Ledenjača u Ćudinoj uvali cave ( WK 46), Veliki Javornik, 44.84° N, 15.57° E (Novak 2004b); Maksimova pećina cave, Jankovac, 657 m, 45.52° N, 17.68° E (Babić 1916); 2 ♁♁, Medveđak Mt., Plitvice, 889 m, 44.89° N, 15.64° E (Novak 2004b); 2 ♁♁, 1 ♀, E Sveti Jakob, stream valley of river Medvešćak, 600 m, under stones and wet logs near river, 45.884° N, 15.945° E, leg. 9 Sep. 2009 ( AXLS 345); 1 juv., near Vrilo Rudnice, Kamenica Skradnička, 260 m, 45.23° N, 15.35° E, leg. Jul. 1987 (Novak 2004b); 1 juv., Ozalj, Karlovac, Kordun, Ozaljska spilja cave, 45.62° N, 15.45° E, leg. 6378 (Novak 2004b); 13 juv., Pećina pri Tržiću (=Pećina kod mlina), Tržić Tounjski, beside the Mrežnica River, 45.23° N, 15.41° E, leg. Jul. 1983 (Novak 2004b); 1 juv., Plitvice (coordinates inaccurate), 677 m, 44.88° N, 15.62° E (Babić 1916); 6 ♁♁, Sagorije, Ogulinsko, Ivancij, hunter house at Smitowo jezero, 351 m, at large rocks in Fagus forest, 45.188° N, 15.223° E, leg. 7 Sep. 2009 ( AXLS 319); 1 ♁, 1 ♀, Sagorije, Ogulinsko, Ivancij, Sagorska peč, 326 m, at wall in cave entrance, 45.197° N, 15.222° E, leg. 7 Sep. 2009 ( AXLS 322); 1 ♁, Skrad, 700 m, 45.43° N, 14.91° E, leg. May 1905 (Babić 1916); 1 juv., Sveta gora, 580 m, 45.53° N, 14.67° E (Novak 2004b, Babić det. as O. alpinus ); 1 ♁, 1 ♀, Vilinska jama cave (=Vilina jama, Volina-Volinjska jama, one hour away from Krapina), leg. 11 Aug. 1885 (Jurinac 1886, Novak 2004b), orig. det. as Liobunum religiosum ).

CZECH REPUBLIC: All records originate from Šilhavý (1981) so far not otherwise indicated. Šumava, Boubín, 48.94° N, 13.79° E; Holná (maybe wrong), 49.12° N, 14.87° E; Horní Smrčné, 49.33° N, 15.76° E; Hrubá Skála, 50.54° N, 15.19° E; Ještěd Mt., 50.73° N, 14.98° E; Jevany, 49.97° N, 14.81° E (Šilhavý did not discriminate betw. L. rupestre or L. tisciae for this record); 2 ♁♁, 1 ♀, limestone caves near Brünn (Brno), leg. 17 Sep. 1971 ( CJM 982); Klučky (mountain ridge near Nový Bor), 50.78° N, 14.50° E; Koněšín, 49.19° N, 16.04° E; Kunratice, 50.01° N, 14.48° E (Šilhavý did not discriminate betw. L. rupestre or L. tisciae for this record); Kvilda, 49.02° N, 13.58° E; Lamberk (Náměšť nad Oslavou), 49.17° N, 16.18° E; Máchovo jezero (Dosky), 50.58° N, 14.67° E; Mariánské Lázně, 49.96° N, 12.70° E; Mohelno (Třebíč), 49.11° N, 16.19° E; Moravský kras, 49.31° N, 16.70° E; Olší (Jindřichův Hradec), 49.16° N, 15.37° E; Pravčická brána (Hřensko), 50.88° N, 14.28° E; Rokytnice nad Jizerou, 50.73° N, 15.45° E; Rýchory, 50.67° N, 15.87° E; Špindleruv Mlýn, 50.73° N, 15.61° E; Tiské skály (Děčín), 50.80° N, 14.04° E; Tremšín (Rožmitál), 49.57° N, 13.78° E (Šilhavý did not discriminate betw. L. rupestre or L. tisciae for this record); Vlčí kopec, 49.15° N, 16.45° E.

GERMANY: Baden-Württemberg, 2 ♀♀, Nature Reserve Wutachschlucht, Stigelesfels, 48.0166° N, 8.9333° E, leg. 1 Oct. 1980 ( CJM 1954); 4 ♁♁, Lotenbachklamm, rock face, 47.85° N, 8.3° E, leg. 25 Jul. 1964 ( CJM 123); 1 ♁, Schwäbische Alb, Höllochschacht bei Dettingen, 48.513° N, 9.346° E, leg. 2.12.1967 ( CJM 1198); 2 ♁♁, Tiefental, Kreis Nürtingen, Gutenberg, Gussmannshöhle, 680 m, 48.543° N, 9.520° E, leg. 18 Feb. 1962 ( SMF 13067/2, Dobat 1963); 2 ♁♁, 11 ♀♀, Schwarzwald, Triberg, 48.13° N, 8.23° E, leg. Aug. 1959 ( SMF 11173/12); 5 ♁♁, 3 ♀♀, S-Schwarzwald, Schwarzatal S Schluchsee, Schwarzahalden, 600 m, Bannwald, 47.74° N, 8.19° E, leg. 14 Sep. 2008 ( CJM 6207); 2 ♁♁, 1 juv., Schwarzwald, Titisee, 865 m, 47.89° N, 8.14° E, leg. 29 Sep. 2013 ( AXLS 1626); Wolfsbrunn near Heidelberg, 49.412° N, 8.746° E (C. Koch 1871; Le Roi 1914); Bayern, Benediktenwand, 47.65° N, 11.47° E (L. Koch 1861); Berchtesgaden, Jenner, 800 m, forest, 47.58° N, 13.00° E (Hammelbacher 1986); 1 ♁, 1 ♀, Ramsau / Wachterl, 1000 m, mixed forest, 47.65° N, 12.866° E, leg. 11 Jul. 1986 ( CJM 2804); 3 ♁♁, Tegernsee, ascend to Wallberg, 700–1000 m, coniferous forest, at cliff, 47.68° N, 11.75° E, leg. 31 Sep. 2006 ( CJM 5318); near Regensburg (GPS for the city), 48.97° N, 12.13° E (Martens 1978); 4 ♁♁, Bavarian Forest, N Finsternau, Siebensteinkopf, 1130–1200 m, 48.95° N, 13.566° E, leg. 23 Aug. 1992 ( CJM 4358); 2 ♀♀, Arber, in the cirque-wall (Karwand), 49.10° N, 13.12° E, leg. 19 Aug. 1964 ( CJM 136); 20 juv., Lusen, Arber, Riesloch Falls, 48.933° N, 13.50° E, leg. 14–17 Aug. 1964 ( CJM 143); 1 ♁, 6 juv., Rachelsee, under stones, 48.975° N, 13.402° E, leg. 16 Aug. 1964 ( CJM 113, CJM 109); 6 ♁♁, Riesloch-Fälle near Bodenmais, 49.05° N, 13.083° E, leg. 18 Aug. 1964 ( CJM 101); 1 ♁, 1 ♀, Garmisch-Partenkirchen, Partnachklamm, 850 m, 47.466° N, 11.116° E, leg. 14 Oct. 1978 ( CJM 1856); 1 ♀, NSG Murnauer Moos, 47.64° N, 11.17° E, leg. 1977/78 ( SMF 33105, Löser et al. 1982); Nürnberg, betw. Bruck and Happburg (coordinates inaccurate), 49.52° N, 11.18° E (L. Koch 1877, as L. bicolor ); Pottenstein, Fränkische Schweiz, 49.77° N, 11.40° E, leg. 1 Oct. 1948 ( SMF 59294, Schönhofer 2005); 2 ♁♁, Waldnaabtal, Oberfranken, 49.6° N, 12.133° E, leg. 22 Aug. 1964 ( CJM 86). Hessen, 1 ♁, Werra-Meissner-Kreis, Meissner, stone cellar near Neuen Schwalbenthaler Erbstollen (No. 4725/58), 580 m, Mittlerer Buntsandstein, 51.2056° N, 9.87 09° E, leg. 25 Oct. 2005 ( CJM 4802, S. Zänker, WK 211); Thüringen, 2 ♁♁, Schwarza Valley near Schwarzburg, 50.633° N, 11.183° E, leg. 28 Sep. 2006 ( CJM 5392); Sachsen, 1 ♀, Erzgebirge (Ore Mountains), at Flossgraben betw. Aue and Bockau, 50.55° N, 12.67° E, leg. 21 Nov. 1926 (Büttner 1930); Lausitzer Mts., Neisse Valley betw. Östritz and Hirschfelde, 50.97° N, 14.92° E (Hiebsch 1972); 8 ♁♁, 1 ♀, S Königstein, stream valley before junction to Cunnersdorf, Biela Stream, 150 m, at stone bridge, 50.903° N, 14.064° E, leg. 9 Sep. 2006 ( CJM 5293); 7 ♁♁, Schöna, near border to Czech Republic, 100 m, coniferous and mixed forest, at night at wood and stones, 50.867° N, 14.217° E, leg. 13 Oct. 2005 ( CJM 4752); 2 ♀♀, same data ( CJM 4753).

HUNGARY: Budapest, Mecsek Mts. (coordinates inaccurate), 47.51° N, 19.01° E (Kolosváry 1929, 1964); 10 ex., Geschriebenstein Nature Park, beside road to Velem (St. Veit), NNE Óház, WSW Köszeg (Güns), 400 m, scree in a fresh deciduous forest with maple, linden, hombeam and beech, 47.22° N, 16.30° E (Komposch 2004); Köszegi-hegység, Köszeg, Hét-forrás, 47.39° N, 16.53° E, leg. 31 Aug. 2005 (Lengyel & Muranyi 2006); Köszegi-hegység, Velem, Szerdahelyi-patak-part, 47.345° N, 16.514° E, leg. 17 Sep. 2005 (Lengyel & Muranyi 2006).

ITALY: Carnia, 46.37° N, 13.14° E (Caporiacco 1922); 1 ♁, Friuli-Venezia Giulia, Prov. Udine, Carnic Alps near Tarvisio, 800 m, 46.50° N, 13.57° E, leg. 1 Oct. 1975 ( CJM 1456); 2 juv., Moggio Udinese, Stavoli Cuel Lung alto, Mt. Zovet ( UM 53), 700 m, Fagetum, pitfall traps, 46.48° N, 13.19° E, leg. 16 Jul. –20 Aug. 2001 ( MFSNU, Novak 2004a); 1 ♁, SE Arta Therme, path in forest, 46.478° N, 13.069° E, leg. 1 Sep. 2012 ( AXLS 1349); 1 ♀, Resia, above Clen, between Casera Coot and Slatina superiore ( UM 73), 1000 m, Fagetum, pitfall traps, leg. 20 Aug.–26 Sep. 2001 ( MFSNU, Novak 2004a); 3 juv., Resia, between Berdo di Sopra and Rio Malidul ( UM 73), 1260 m, 46.34° N, 13.29° E, leg. 12 Jun. 2003 ( MFSNU, Novak 2004a); Südtirol, Meran, 46.68° N, 11.17° E (L.Koch 1869, type locality of Leiobunum glabrum L. Koch, 1869 ).

POLAND: 3 ♁♁, Kitz....? (maybe Kitzelberg; label not legible) Bober Katzbach Mts. (GPS not accurate), 50.96° N, 15.84° E, leg. 6 Sep. 1924 ( ZMB 12643). Karkonosze Mts. (Staręga 2004).

SERBIA: 1 ♀, Valjevo, planina Medvenik, 44.24° N, 19.63° E, leg. 1 Aug. 1998 ( CIK 1337); 1 ♁, Valjevo, planina Medvenik, Drenaička pecina, leg. 20 Oct. 1999 ( CIK 1336).

SLOVENIA: Alto Isonzo, Monte Merzlì, 46.22° N, 13.70° E, leg. Nov. 1915 (Marcellino 1973); 1 ♁, Celje (Cilli), deciduous forest, 46.24° N, 15.26° E, leg. 25 Oct. ( ZMB 12635); 7 ♁♁, Huda Luknaja Cave, Tisnick Mt. SE Slowenj Gradec, 46.48° N, 15.07° E, leg. 16 Oct. 2004 ( CJM 4203); 3 ♀♀, Huda Luknaja Cave, Tisnick Mt., SE Slowenj Gradec (GPS S of city), 46.48° N, 15.07° E, leg. 16 Oct. 2004 ( CJM 4204); 15 ♁♁, 4 ♀♀, Julian Alps, road to Mount Mangert, upper end of village Strmec na Predelu, 1013 m, 46.41769° N, 13.60833° E, leg. 28 Aug. 2009 ( AXLS 264); 4 ♁♁, Karawanken, road to Loibl Pass, 650 m, 46.40° N, 14.28° E, leg. 2 Oct. 1975 ( CJM 1445); 6 ♁♁, 1 ♀, Ljubljana, small forest at northern city limit, 46.10530556° N, 14.51822° E, leg. 3–4 Sep. 2012 ( AXLS 1389); 9 ♁♁, 1 ♀, Pohorje Mts., Mislinja, locality Glažuta, 1013 m, stream valley, on cliff-faces in coniferous forest, 46.480° N, 15.280° E, leg. 26 Aug. 2009 ( AXLS 237); 3 ♁♁, Savinje valley, Erjavčeva Jama, entrance zone, 700 m, 46.383° N, 14.717° E, leg. 2 Oct. 1994 ( CJM 3161); 1 ♀, Slowenj Gradec, at housewall, 46.51° N, 15.07° E, leg. 13–17 Oct. 2004 ( CJM 4140); Triglav, Martuljuk, 46.46° N, 13.83° E, leg. 15 Aug. 1928 (Hadži 1931); 1 juv., vicinity of Zirknitz (= Cerknica), 650 m, Fagus forest, 45.79° N, 14.37° E, leg. 17 Jul. 1974 ( CJM 1260); 3 ♁♁, 1 ♀, Unec, Rakov Škocjan, at border of river Rak, 520 m, at cliff in Fagus forest, leg., 45.796° N, 14.293° E, leg. 10 Sep. 2009 ( AXLS 352); 1 ♁, Wochein (= Bohinj, Julian Alps) ( ZMB 12637).

SWITZERLAND: Basel-Landschaft, Arlesheim, 47.49° N, 7.63° E (Müller & Schenkel 1895; Lessert 1917).