Zetemenos sexlineatus Bode, 1953

Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong & Prokop, Jakub, 2012, Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga), Acta Entomologica Musei Nationalis Pragae 52 (1), pp. 89-127 : 124

† Zetemenos sexlineatus Bode, 1953

( Figs. 93 , 119 )

Zetemenos sexlineatus Bode, 1953: 230 . Zetemenos sexlineatus: PONOMARENKO (1992: 186) .

Type locality and age. Germany, Niedersachsen, Grassel at Braunschweig (current GPS coordinates: 52.4°N 10.6°E. Posidonia Shale Formation, Early Jurassic, Early Toarcian, 183.0–182.0 mya ( CLAPHAM 2011b).

Material examined. HOLOTYPE: SMF VI 1399 (piece only).

Taxonomic note. The holotype, which is the only available specimen of this species, is a badly preserved fossil in dorsal view showing an elongate beetle with striate elytra. Other characters are not preserved, except for the abdomen with five ventrites, narrow and elongate metepimera, and possibly the frontoclypeal suture (however, the impression on the head may also represent some ventral head structures). The latter character would be also the only possible indication of the position of the species in the Hydrophiloidea . However, due to the bad preservation we do not consider this character as adequate to justify the superfamily assignment of the fossil. For that reason, the species including the currently monotypic genus Zetemenos Bode, 1953 to which it belongs are excluded from the Hydrophiloidea and transferred to Coleoptera incertae sedis.

Figures

Figs. 89–93. Representatives of Helophorus, subgenus Mesohelophorus Ponomarenko, 1977 (89–90) and taxa excluded from the helophorid lineage (91–93). 89–90 – Helophorus palaeosibiricus nom. nov., PIN 3063/841, holotype, piece and counterpiece; 91 – Helophoropsis brodiei (Giebel, 1856), holotype. 92 – Tychon antiquum (Giebel, 1856). holotype. 93 – Zetemenos sexlineatus Bode, 1953, holotype. Scale bars: 0.5 mm.

Figs.113–119.Taxa excluded from the helophorid lineage. 113 – Helo- phoropsis brodiei (Giebel, 1856), holotype; 114 – ‘Mesosperchus’ angulatus Ponomarenko, 1985, holotype; 115 – ‘Mesosperchus’ notatus Ponomarenko, 1977, holotype; 116 – Tychon antiquum (Giebel, 1856), holotype; 117–118 – ‘Mesosperchus’ schultzi Ponomarenko, 1985, holotype, piece and counterpiece; 119 – Zetemenos sexlineatus Bode, 1953, holotype. Scale bars: 1 mm.