Laetopsia shatrovskiyi ( Prokin, Ren & Fikáček, 2010 ) Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop, 2012

Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong & Prokop, Jakub, 2012, Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga), Acta Entomologica Musei Nationalis Pragae 52 (1), pp. 89-127 : 98

† Laetopsia shatrovskiyi ( Prokin, Ren & Fikáček, 2010) comb. nov.

( Figs. 12–13 , 49–52 )

Hydrophilopsia shatrovskiyi Prokin, Ren & Fikáček, 2010: 178 .

Type locality and age. China, Liaoning Province, Shangyuan County, Chaomidian Village, Huangbanjigou [the name of the villige was misspelled in the original descirption]. Yixian Formation, Jurassic-Cretaceous boundary, Late Tithonian-Berriasian, ca. 145–140 mya ( REN et al. 2010). An alternative dating was proposed by SWISHER et al. (1999): Lower Cretaceous, Aptian, 124.6 mya (see REN et al. (2010) for discussion on various datings of Yixian formation).

Material examined (3 spec.). HOLOTYPE: CNU 2010242 (piece only) . PARATYPES: CNU 2010016 + CNU 2009199 (piece and counterpiece) ; CNU 2010012 (piece only) .

Redescription. Body elongate oval, body 5.0– 5.5 mm long (holotype 5.4 mm). Head with protruding eyes, clypeus subquadrate, with labrum attached to its anterior margin; frontoclypeal suture distinct, Y-shaped, slightly grooved medially; head surface with weak granulation. Mentum slightly wider than long, gula slightly constricted but with well-separated gular sutures. Antennae with three-segmented antennal club. Pronotum arcuate laterally, pronotal flanks narrow, narrowing posteriad. Prosternum long anterior to procoxae. Mesanepisterna probably nearly connecting anteriorly, mesoventrite extremely narrow at anterior margin; posterior portion of mesoventrite with very distinct transverse ridge. Mesocoxae large, contiguous. Elytron with striae 1–7 darkened; epipleuron narrow, subdivided into inner and outer portion, inner portion narrow. Metaventrite longer than mesoventrite, metepimeron with oblique transverse ridge anteriorly. Metacoxae transverse, abdomen with 5 ventrites. Legs rather long, tibiae slightly longer than femora, tibial apices with strong spines.

Note. One paratype of this species (CNU2010004) was found not to be conspecific with the remaining specimens and is described below as Helophorus (Mesosperchus) yixianus sp. nov.

Figures

Figs.9–13. Laetopsia gen. nov. 9 – L. hydraenoides (Prokin, Ren & Fikáček, 2010), holotype. 10–11 – L. mongolica (Ponomarenko, 1987) (10 – PIN 3664/1567, holotype; 11 – PIN 3664/1661). 12–13 – L. shatrovskiyi (Prokin, Ren & Fikáček, 2010) (12 – CNU 2010199; 13 – CNU 2010242, holotype). Scale bars: 2 mm (a: Figs. 9, 12–13; b: Figs. 10–11). Abbreviations: ant – antenna; sct – scutellum.

Figs. 42–52. Representatives of Hydrophilopsia Ponomarenko, 1986 and Laetopsia gen. nov. 42–48 – Hydrophilopsia longitarsalis Ponomarenko, 1986, holotype (42–43 – piece and couterpiece, general views; 44 – detail of the head and thorax, photo under alcohol; 45 – same, photo in dry state; 46 – detail of aedeagus; 47 – legs of piece, 48 – legs of counterpiece). 49–52 – Laetopsia shatrovskiyi (Prokin, Ren & Fikáček, 2010) (49, 51 – CNU 2009199; 50, 52 – CNU 2010242).