Campanula versicolor Andrews (1804

Janković, Ivana, Lakušić, Dmitar, Pietro, Romeo Di & Kuzmanović, Nevena, 2017, Nomenclatural notes and typifications in Campanula versicolor (Campanulaceae) and related names, Phytotaxa 323 (3), pp. 264-274 : 265-267

Campanula versicolor Andrews (1804 : t. 396)

Lectotype (designated here):—ILLUSTRATION. plate CCCXCVI [tab. 396] in Andrews, Bot. Repos. 6: t. 396 (1804)(available at http:// plantillustrations.org/illustration.php?id_illustration=111483&mobile=0&code_category_taxon=1)( Fig. 1A ).

Epitype (designated here):— GREECE. Parnas [Mt Parnassus], stene, krečnjak [limestone cliffs], 38.56761° N, 22.56891° E, 1564.2 m a.s.l., 21 August 2014, D. Lakušić, N. Kuzmanović & I. Janković 40045 (BEOU!)( Fig. 2 ).

Protologue information:—“It was first raised from seeds brought to this country from Greece, by the late and much regretted Professor Sibthorp”.

Note: —The color illustration is published in the protologue together with the description, being part of original material. According to Stafleu & Cowan (1976), the drawings in the Botanist‘s repository are made by Andrews based on the living specimens, and no herbarium material is known to exist for any of his drawings. The plant was raised from the seeds brought from Greece by Prof. Sibthorp ( Andrews 1804). The description was made based on the living specimen as well. However, this species was discovered by John Sibthorp during his first field trip to the Mediterranean regions— Greece and Asia Minor in 1786–1787. Unfortunately, he died soon after the return from the second field trip made in between 1794–1795, before he was able to publish the large part of his work from the Mediterranean field trips, intended to be the part of his Flora graeca. During this first field trip, Sibthorp was accompanied by the artist Ferdinand Bauer, who made the botanical illustration including the one of C. versicolor ( Fig. 1B )( Stafleu & Cowan 1985). The task of preparing and publishing Sibthorp‘s work was undertaken by J. Smith, who issued the first volume of „Flora Graeca prodromus“ two years after Andrews‘ publication. In that book, a short description of C. versicolor is also provided. However, Campanula versicolor Sibthorp (in Sibthorp & Smith 1806: 138) is a later homonym of C. versicolor Andrews and is illegitimate according to Art. 53 of the ICN.

In addition, for the purpose of the precise application of the name, the specimen preserved at the herbarium of the University of Belgrade (BEOU 40045) is designated here as epitype of C. versicolor . The habitus and capsule, as well as the dimension of C. versicolor , is not evident in the lectotype. Thus, an epitype representing the modern concept of the species is designated to support the lectotype. The material was collected on Mt Parnassus ( Greece) which was one of the localities visited by Sibthorp during his expedition on the Balkan Peninsula in 1787 ( Bruce 1970)( Fig. 2 ).

Figures

FIGURE 1. A. Lectotype of Campanula versicolor Andrews. B. Color illustration of C. versicolor Sibthorp.

FIGURE 2. Epitype of Campanula versicolor Andrews (BEOU 40045!).