Amphineurus (Rhamphoneurus) oosterbroeki, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.20. Amphineurus (Rhamphoneurus) oosterbroeki sp. nov.

Material examined.

Holotype: ♂, Chile, Chiloé Is., Chaiten [42°58′S 72°32′W], 5/ 8-II-1954, Peña (USNM) *. Paratypes: 1 ♂, Chile, Chiloé Is., Chaiten [42°58′S 72°32′W], 5/ 8-II-1954, Peña (USNM) *; 1 ♂, Chile, Llanquihue, Carelmapu [41°44′S 73°40′W], Coast of Canal de Chacao , 18-III-1957, Peña (USNM) *; 1 ♂, Chile, Osorno, Bahia Mansa [40°35′S 73°44′W], South of Pucotrihue, 12-IV-1968, Peña (USNM) *.

Etymology.

The species is named after Pjotr Oosterbroek, an important Tipulomorpha expert.

Diagnosis.

This species is distinguished by only a dark spot on R2, basal deflection of R5 as long as r-m, and M1+2 longer than basal deflection of M1. Furthermore, this species is characterized by male terminalia with sheath of aedeagus with a triangular spur and the tip of sheath inclined.

Description.

Wing length 5.47 mm, width 1.68 mm. - Coloration: General coloration brown. Rostrum and palpus brown. First segments of antenna yellow, remainding segments yellowish brown. Head brown. Abdomen yellowish brown. - Head (dorsal view Fig. 25B , ventral view Fig. 25A ): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape narrower than pedicel; terminal flagellomere dilated. - Thorax: Wing (Fig. 25D ) almost clear of markings, except for marking on R2; veins near fork of bM (Fig. 25C ) faded; M1+2 2.5 × longer than basal deflection of M1; R2+3+4 longer than R2+3. - Male terminalia (Fig. 25E ): Male tergite IX slightly darkened along posterior margin, with deep median U-shaped notch. Ventral branch of gonocoxite slender, very long, setose on distal tip. Lobe of gonostylus with rounded lobule; lobule longer than wide, shorter than half of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved. Mesal lobes of gonocoxite asymmetrical; left mesal lobe curved; right mesal lobe short with sharp turn. Sheath of aedeagus straightened, with wide rectangular projection; darkened only near tip.

Remarks.

Specimens of this new species were previously identified by C.P. Alexander as A. (R.) insanus Alexander. This species resembles A. (R.) podenasi sp. nov. but differs mainly in the length of r-m, concentration of setae on gonocoxite, shape of male tergite IX and right mesal lobe.

Figures

Figure 25. Amphineurus (Rhamphoneurus) oosterbroeki sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E Male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; cgonst, clasper of gonostylus; goncx, gonocoxite; lgonst, lobe of gonostylus; ms lobe, mesal lobe of gonocoxite; t 9, male tergite IX.