Laena quadricollis Weise, 1878

Schawaller, Wolfgang, 2020, New taxonomic and faunistic data to species of the genus Laena Dejean from Turkey (Coleoptera, Tenebrionidae, Lagriinae), with descriptions of two new species, Zootaxa 4861 (3), pp. 385-398 : 396

Laena quadricollis Weise, 1878

( Figs. 7 , 44, 45 )

Examined specimens: Russia, Krasnodar, W Maikop, 300 m, 8. VII .1986, leg. S. Golovatch, 4 ex. SMNS .— Georgia, Rike , N Sugidi, 24.IX.1986, leg. S. Golovatch, 1 ex. SMNS .— Adzaria, Batumi , VI.1975, leg. Rous, 1 ex. SMNS .— Georgia, Shida-Kartli, Tkemlovana , 2000 m, 29.V.–28.VI.2016, leg. D. Forminykh, 1 ex. CPE .— Georgia , Letchkhumskiy Mts. , Khideshlebi, 1235 m, 5.VII .2015, leg. A. Pütz, 1 ex. CPE , 1 ex. SMNS .— Georgia, Adjara, 2 km SE Tskhmorisi , 700–900 m, 13.VI.2018, leg. A. Pütz, 2 ex. CPE .— N Ossetia, 1.7 km SE Nizhniy Tsey , 1400 m, 30.V.–12. VII .2016, leg. D. Fominykh, 2 ex. CPE .— S Ossetia , 1 km NW Bakhuta, 1250 m, 28.V.–12. VII .2016, leg. D. Fominykh, 1 ex. CPE .— Ossetia, Ratchinsky Mts. , Kobet, 1800 m, 16. VII .2016, leg. D. Fominykh & A. Zubov, 1 ex. SMNS .— Turkey, Artvin, below Pirnalli, Karkal Daği , 1250 m, 11.VI.1986, leg. C. Besuchet, I. Löbl & D. Burckhardt, 1 ex. SMNS .— Turkey, Artvin, 8 km W Borçka , 350 m, 8.VI.1986, leg. C. Besuchet, I. Löbl & D. Burckhardt, 1 ex. SMNS .— Turkey, 50 km NW Artvin , Murgul, 1300 m, 13. VII .1998, leg. Skoupý, 1 ex. SMNS .— Turkey, Artvin, S Arhavi, 9.–14. VII .1977, collector unknown, 1 ex. MNB .— Turkey, Rize, 25 km S Pazar , 670 m, 11. VII .2008, leg. M. Schülke, 4 ex. MNB .— Turkey, 25 km SE Rize , S Kaptanpasa, 690 m, 2. VIII .2006, leg. V. Assing & M. Schülke, 1 ex. CPE , 1 ex. SMNS .— Turkey, 25 km SE Rize, 7 km E Ikizdere , 1030 m, 31. VII .2006, leg. V. Assing, 1 ex. CPE .— Turkey, 25 km SE Rize, 4 km E Ikizdere , 750 m, 31. VII .2006, leg. V. Assing, 1 ex. SMNS .— Turkey, 40 km SSW Giresun, 1140 m, 29. VII .2006, leg. V. Assing, 1 ex. CPE .— Turkey, 30 km S Giresun, 830 m, 28. VII .2006, leg. V. Assing, 1 ex. CPE .— Turkey, 50 km SW Trapzon , NE Kürtün, 1430 m, 27. VII .2006, leg. M. Schülke, 1 ex. SMNS .— Turkey, Artvin, Cankurtaran Geçidi , 700 m, 1.–3. VII .1996, leg. D. Szalóki, 1 ex. SMNS .— Turkey, Artvin, 20 km W Borçka, Cankurtaran Geçidi , 700 m, 730–775 m, 29.–30.VI.2004, leg. J. Hájek & J. Růžička, 1 ex. SMNS .— Turkey, Artvin, 10 km W Borçka , 800 m, 25.VI.1998, leg. P. Vonička, 1 ex. SMNS .— Turkey, Artvin, Cankurtaran Pass , 800 m, 1.–2. VII .1996, leg. A. Podlussány, 1 ex. SMNS .

Remarks: This species was described from the Caucasus and so far unknown from Turkey.However, the collected specimens in the Black Sea range completely coincide with available specimens from adjacent southwestern Caucasus, also in the shape of the aedeagus( Figs.44,45 ).This distributional pattern in Pontus and Caucasus fits also under zoogeographicalaspects. L.graeca alsofromnorthernTurkeyissomewhatsimilar,buttheaedeagusisdistinctlydifferent( Fig.42 ). Published locality: Zirkassia (Caucasus) ( Schuster 1916).

Figures

FIGURES 1–8. Dorsal view of Laena species from Turkey (and few from Greece and Bulgaria). 1 L. ganglbaueri, Istanbul, SMNS. 2 L. apfelbecki, Greece Évia, SMNS. 3 L. weisei, Greece Sapka, SMNS. 4 L. kraatzi, Sinop, SMNS. 5 L. osmanlis, Bursa, SMNS. 6 L. assingi sp. nov., paratype SMNS. 7 L. quadricollis, Trabzon, SMNS. 8 L. theana, Isparta, SMNS. Scale line: 3.0 mm.

FIGURES 22–46. Right posterior male tibia of Laena species from Turkey and Greece (Figs. 22–25), and apicale of aedeagus of Laena species from Turkey and few from Greece (Figs. 26–46). 22 L. apfelbecki, Greece Évia, SMNS. 23 L. bozdagica sp. nov., holotype Izmir, SMNS. 24 L. kraatzi, Greece Chios, SMNS. 25 L. lindbergi, Bingöl, SMNS. 26 L. deplanata, Izmir, SMNS. 27 L. ferruginea, Greece Corfu, SMNS. 28 L. ferruginea, Antalya, SMNS (det. Kaszab as L. liliputana syn. nov.). 29 L. wanensis, Nur Daglari, SMNS. 30 L. bozdagica sp. nov., paratype Izmir, SMNS. 31 L. merkli, Hakkari, CPE. 32 L. obscuricornis, Meselik, SMNS. 33 L. byzantina, Artvin, SMNS. 34 L. korbi, Konya, SMNS. 35 L. lindbergi, Bitlis, SMNS. 36 L. apfelbecki, Greece Evia, SMNS. 37 L. pici, Antakya, SMNS. 38 L. ganglbaueri, Alem Dagh, SMNS. 39 L. weisei, Greece Sapka, SMNS. 40 L. theana, Erfelek, SMNS. 41 L. assingi sp. nov., holotype Fethiye, CPE. 42 L. graeca, Bolu, SMNS. 43 L. osmanlis, Bursa, SMNS. 44 L. quadricollis, Rize, SMNS. 45 L. quadricollis, Krasnodar Maikop, SMNS. 46 L. kraatzi, Alem Dagh, SMNS. Scale line: 1 mm.