Andraegoidus richteri ( Bruch, 1908 )

R, Juan Pablo Botero & Monné, Marcela L., 2012, Revision of the genus Andraegoidus Aurivillius (Insecta: Coleoptera: Cerambycidae), Zootaxa 3169 (1), pp. 1-60 : 34-39

Andraegoidus richteri ( Bruch, 1908) status nov.

( Figs 131–156 , 228 )

Trachyderes richteri Bruch, 1908: 215 ; 1912: 205 (cat.); Aurivillius, 1912: 480 (cat.); Bruch, 1912: 205 (cat.); Schwarzer, 1929: 361, fig. 13; Blackwelder, 1946: 591 (cat.); Bachmann & Di Iorio, 2002: 77.

Andraegoidus rufipes richteri ; Hüdepohl, 1985: 58, fig. 28; Monné, 1994: 87 (cat.); Monné & Giesbert, 1995: 142 (check); Monné, 2005: 583 (cat.); Monné & Bezark, 2010: 151 (check).

Male. Integument from black to brown. Antennomeres III–XII orange with apex black or dark brown. Elytra bicolor: brown with apex black or totally black or light brown.

Antennal tubercles rugose and with coarse punctures. Upper eye lobes well-separated, distance between each upper lobe equal to twice the width of an upper lobe. Submentum excavated. Genae with dense and moderately coarse punctures, short, decumbent, and whitish hairs. Antennae 12–segmented, exceeding elytral apices, maximum at segment IX. Scape strongly punctate, expanded at apical region; with short and sparse hairs mainly concentrates in ventral region. Pedicel with pubescence short and dense; segment III with pubescence short and dense and uniform punctures, except at apex which is smooth; segments IV–XII smooth. Segment III 1/5 longer than scape and IV, segments IV–XI subequal in length, segment XII 1/4 shorter than XI and slightly curved.

Prothorax about 2/5 wider than long; coarse and dense sexually dimorphic punctation; surface with aspect alveolate, feebly demarcated ( Fig. 131 ). Antemedian tubercle visible and elevated, smaller than postmedian, postmedian tubercle at right angle to axis of body. Pronotum with sexually dimorphic punctation concentrated principally in anterior and lateral region, absent on posterior region, transverse carina, median and postmedian elevations, anterior and posterior depressions and fine median longitudinal band in anterior region; elevations may possess coarse and sparse punctures. Sexually dimorphic punctation of pronotum with indentations shallow, rounded and ovoid ( Fig. 132 ), denser and elongated at lateral region; each indentation with a single or with a several distinct cluster of pores ( Fig. 133 ), distributed in all indentation; between indentations are scattered pores that may possess a short seta which is generally at apex ( Fig. 134 ). Prosternal sulcus with dense, long and whitish hairs. Prosternal process smooth elevated and slightly narrowed at middle, at most 1/3 as broad as the procoxal cavity. Mesosternal process, at most subequal as broad as mesocoxal cavity; with discrete projection directed anteriorly; posterior margin sinuous.

Scutellum approximately 1/6 length of elytra, with fine punctures. Elytra at least in anterior ¾ subparallel; three times longer than prothorax. Surface smooth or with some fine punctures on anterior region. Apices variables: curved, slightly truncate; unarmed.

Sternites I–IV with long, fine, and whitish hairs. Sternite V covered with dense and short pubescence; posterior margin truncate or slightly sinuous. Sternite I 2.5 times longer than II; sternites II–IV gradually decreasing in length; V subequal in length to IV.

Terminalia: Tergite VIII ( Fig. 140 ) with apical margin adorned with yellow hairs. Sternite VIII ( Fig. 139 ) transverse, apical margin emarginated and with long hairs at side and decreasing in length to middle which is glabrous; sternal apophysis about half as long as sternite. Ventral arc ( Fig. 141 ) with short apophysis, about 1/10 as long as arms. Dorsal arc ( Fig. 142 ) “U” shaped; apical margin with some sparse hairs. Tegmen ( Figs 143–145 ) ¼ shorter than median lobe; distal region divided into parameres, which are cylindrical-elongate, narrowed to apex and with long hairs; ring piece elongated, fused at apex, with short apical projection. Median lobe ( Figs 146–148 ): dorsal lobe slightly shorter than ventral lobe; apex of dorsal lobe rounded and ventral lobe acuminate; basal apophysis ⅓ longer than apical region. Internal sac with sclerotized piece “C” shaped.

Female. Antennae shorter than in males, exceeding elytral apices at segment XI. Prothorax smooth or with fine and sparse punctures; less swollen than in males, lateral tubercles more visible, antemedian tubercle subequal to postmedian tubercle. Transverse carina and elevations with or without coarse and dense punctures, with long and sparse hairs, denser in region of lateral tubercles; postmedian elevations more elevated than carina and with rounded top. Lateral regions of prosternum with microsculptured surface and with dense wrinkles ( Fig. 156 ).

Terminalia: Tergite VIII ( Fig. 149 ) with apical margin sinuous. Posterior region of sternite VIII ( Figs 135–138 ) two times wider than long; with a row of aciculate setae as long as conchoidal setae, conchoidal and petiolate setae longer and curved at apex, conchoidal setae with surface moderately wavy and with some slits; petiolate setae with dorsal slits; sternal apophysis about 2/5 longer than posterior region. Ovipositor ( Figs 150–151 ) with short and parallel lateral lobes, margin with long, fine and sparse hairs, inner margin straight near vulva.

Measurements (mm). female/male. n= 4/8. Total length, 22.05±4.02/20.53±5.13; prothorax length, 5.10±0.89/4.65±1.21; greatest prothorax width, 7.70±1.76/6.73±1.92; elytral length, 15.05±2.83/13.65±3.67; humeral width, 7.55±1.65/6.84±1.90.

Type material. We examined the female paratype ( Fig. 152 ), from Catamarca, Argentina (MZSP) .

Discussion. Hüdepohl (1985) recognized Trachyderes richteri as a subspecies of Andraegoidus rufipes and characterized it by having the prothorax with coarse sexually dimorphic punctation and coarse punctures in the postmedian elevations of the pronotum. Through examination of the female paratype of T. richteri ; lectotype of T. ebeninus ; type-specimen photos from some of the synonyms of A. rufipes rufipes ( T. rufipes var. axillaris , T. rufipes var. brunnipennis , T. boisduvalii ) and approximately 400 specimens of A. rufipes rufipes and A. rufipes richteri , additional significant differences were discovered in A. rufipes richteri : elytra subparallel at least in anterior ¾.; lateral regions of prosternum with microsculptured surface and with dense wrinkles ( Fig. 156 ). Andraegoidus rufipes possesses a prothorax with fine sexually dimorphic punctation; smooth postmedian elevations; elytra gradually decreasing to apex; females with smooth lateral regions of prosternum. Based on the examined material examined and these additional morphological differences, we propose the elevation of this species to A. richteri .

Geographical distribution. Bolivia (LP), Argentina (CA; JU; SA; TU). A new country record is added from Paraguay (“Región del chaco”) ( Fig. 228 ).

Specimens examined. BOLIVIA, La Paz: male, 1947 ( MNRJ) . PARAGUAY, “ Región del Chaco ”, male ( MNRJ) . ARGENTINA , Catamarca: male ( MNRJ) ; female (Paratype), “190?”, Bruch leg. ( MZSP) ; male, 30.XI.1907, Bruch C. leg. ( MNRJ) ; Jujuy: female ( MNRJ) ; Salta: Amblayo , male, 7.II.1945 ( MNRJ) ; Tucumán: female, I.1949 ( MNRJ) ; Trancas, San Pedro de Colalao ; female and male, I.1949; male, II.1950; male, II.1953, Arnau leg. ( MNRJ) .

Figures

FIGURE 228. Map of geographical distribution of Andraegoidus fabricii, Andraegoidus homoplatus, Andraegoidus richteri, Andraegoidus rufipes.

FIGURES 131–138. Andraegoidus richteri. 131–134 pronotum, male: 131, general view; 132, detail of sexual punctation; 133, detail of indentation with pores; 134, detail of a point with setae; 135–138, sternite VIII, female: 135, general view; 136, detail of brush of setae; 137, detail of conchoidal setae; 138, detail of petiolate setae.

FIGURES 139–148. Andraegoidus richteri male. 139, sternite VIII; 140, tergite VIII; 141, ventral arc; 142, dorsal arc; 143– 145, tegmen: 143, dorsal view, 144, lateral view, 145, ventral view; 146–148, median lobe: 146, dorsal view, 147, lateral view, 148, ventral view.

FIGURES 149–151. Andraegoidus richteri female. 149, tergite VIII; 150–151, ovipositor: 150, dorsal view, 151, ventral view.

FIGURES 152–163. 152, Andraegoidus richteri, paratype, female; 153–154, Andraegoidus richteri, male; 155, Andraegoidus richteri, female; 156, detail of head and thorax, lateral view, female; 157, Andraegoidus fabricii, neotype, male; 158–161, Andraegoidus fabricii, male; 162–163, Andraegoidus fabricii, female.