Guyotemaimetsha enigmatica Perrichot, Nel & Neraudeau , 2004a

Perrichot, Vincent, Ortega-Blanco, Jaime, McKellar, Ryan C., Delclos, Xavier, Azar, Dany, Nel, Andre, Tafforeau, Paul & Engel, Michael S., 2011, New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae), ZooKeys 130, pp. 421-453 : 426-429

Guyotemaimetsha enigmatica Perrichot, Nel & Neraudeau, 2004a Figs 14

Guyotemaimetsha enigmatica Perrichot, Nel & Néraudeau, 2004a: 159, figs. 1A-B, 2 [♂].

Type material.

Holotype MNHN.F.A30174 (♂); paratype MNHN.F.A30175 (♂); in amber collection of the Museum National d’Histoire Naturelle, Paris, France.

Other material examined.

Specimens IGR.ARC-1 (♀, missing anterior part of the head and with cuticle somewhat altered by taphonomy), IGR.ARC-309.11 (sex unknown, missing apices of right forewing and metasoma), IGR.ARC-370.7 (= scan ESRF-A-018g, reconstructed, ♀, in opaque amber), IGR.ARC-378.1 (♂, missing head, most metasoma and apices of all legs), IGR.ARC-378.2 (scan ERSF A-401-378, not reconstructed; ♂), IGR.ARC-393.4 (= radiograph ESRF-A-053d, not reconstructed; ♂), IGR.CDL-2.5 (♂, with forewing fragment in IGR.CDL-2.16, missing apical flagellomeres of left antenna; ex collection Arnaud), IGR.CDL-2.33 (♂, missing left antenna and apices of right antenna, right forewing, and all legs; ex collection Arnaud). All specimens deposited in the amber collection of the Department of Geosciences of the University Rennes I, France.

Indeterminate specimens.

IGR.ARC-171 (♂, missing head, anterodorsal part of metasoma, and forelegs); IGR.ARC-191 (♂?, missing part of the head and mostly obscured, preserved in very dark amber); IGR.ARC-275 (♀, missing parts of mesosoma, left legs, and with metasoma mostly obscured by a bacterial film). These specimens agree in most respects with the definition of Guyotemaimetsha enigmatica but little is preserved and attribution to this species cannot be asserted. All specimens deposited in the amber collection of the Department of Geosciences of the University Rennes I, France.

Type locality.

Font-de-Benon quarry, 1 km east of Archingeay, Charente-Maritime, France.

Stratigraphic horizon.

Lithological subunit A1sl-A, Uppermost Albian–Lowermost Cenomanian, middle Cretaceous.

Diagnosis.

As for the genus (see above).

Redescription.

Male: Integument reddish-brown in color, with dense, minute punctures and fine, decumbent pubescence. Head transverse, with length approximately 0.8x width; ocelli separated from each other by one ocellar diameter; antennae inserted between compound eyes, closer to each other than to compound eye margin or posterior clypeal margin; scape globular in lateral view, laterally compressed in frontal view, with faintly concave apex; pedicel with sides convex; flagellum with dense coat of short, inclined setae, with first article longest, twice length of pedicel; penultimate article as broad as long; apical article with rounded apex; clypeus transverse, about twice as broad as long, with anterior margin rounded and posterior margin straight; mandibles large, with outer margin convex, overlapping apically when closed; right mandible with basal tooth large, second and third teeth equally smaller, apical tooth largest; left mandible with basal tooth large, median tooth smallest, apical tooth largest. Mesosoma compact, 0.7x as high as long; pronotum in dorsal view reduced to a short neck; mesoscutum broader than long, about 0.4x mesosomal length, abruptely truncate anteriorly to form a flat vertical surface, bordered by a transverse carina at anterolateral corners, with low dorsal convexity; median mesoscutal sulcus crenulate, notauli deeply impressed, slightly diverging anteriorly; mesoscutellum with low dorsal convexity, small triangular axillae separated anteriorly by large foveate groove; propodeum areolate, gradually sloping to pronounced posterior lip; meso- and metapleuro n with anterior margin foveolate. Forewing hyaline, with costal cell distinctly enlarged around midlength; Rs junction with Sc+R forming an acute angle; cell [1M] with vein M slightly arched, roughly aligned with Rs, vein Cu shortest, M and Rs+M subequal in length; cells [1M] and [1Rs] well distant from each other, separated by long vein 2Rs+M; 2Cu significantly longer than 1Cu and 1m-cu; M and Cu nebulous at apex, reaching wing margin; 2A fading to nebulous vein posterior of [2Cu]. Hind wing with Sc+R, R, M+Cu, Cu, and A tubular, other veins nebulous; Sc+R and R more heavily sclerotized than M+Cu, Cu, and A; Cu fading to nebulous vein apically; free apex of Rs and Cu short. Metacoxa approximately 1.4x length of pro-and mesocoxae, all with broad bases and distinct apical constriction, in lateral view inserted posteroventrally on mesosoma; metafemur distinctly enlarged around midlength; all tibiae approximately subequal to femora in length, with basitarsus length slightly less than combined length of all subsequent tarsomeres. Metasoma compact, as long as mesosoma; parameres trapezoidal, narrowed toward apex, with outer surface slightly convex, with short erect setae around apex; cerci small, spatulate, inserted just anterior to parameres.

Female: Very similar to male but with pedicel cylindrical and each leg with a small plantar lobe (or tarsal plantula) on ventral apex of tarsomeres I–IV; ovipositor approximately as long as metasomal length, apex acute. Integument sculpturing and pubescence not visible, altered on specimen IGR.ARC.1 and not reconstructed in microtomographic scan of specimen IGR.ARC-370.7.