Pupina brachysoma Ancey, 1904

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina brachysoma Ancey, 1904

Figs 36M , 41F

Pupina brachysoma Ancey in Bavay & Dautzenberg, 1904 [1903]: 230, 231, pl. 10, figs 15, 16. Type locality: Haut-Tonkin [northern Vietnam]. Fischer-Piette 1950: 171. Wood and Gallichan 2008: 31, pl. 25, figs 5, vi.

Type material examined.

Lectotype MNHN-IM-2000-9652 (Figs 36M , 41F ) from Haut Tonkin. Paralectotypes NMW.1955.158.24153 figured in Wood and Gallichan (2008: pl. 25, figs 5, vi).

Diagnosis.

Shell ovate-fusiform; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened, slightly expanded. Parietal tooth sharp, tooth-like, thickened; columellar tooth fin-shaped, slightly thickened, located next to slit-like anterior canal. Posterior canal widened.

Differential diagnosis.

Pupina brachysoma is most similar to P. tchehelensis and P. lowi in having a sharp, tooth-like, thickened parietal tooth, a fin-shaped, thickened columellar tooth that is located next to a slit-like anterior canal, and a posterior canal that is gradually widening. However, P. brachysoma is different from both P. tchehelensis and P. lowi by a more ovate-fusiform shell shape, and a less thickened but slightly expanded apertural lip. Pupina brachysoma is also similar to P. dorri dorri in shell shape, but differs in having a gradually widening posterior canal.

Distribution.

Known only from the type locality ( Bavay and Dautzenberg 1904).

Remarks.

As the original description did not explicitly state that the description of this species was based on a single specimen (nor could this be inferred), the designation of a holotype by Fischer-Piette (1950) in fact constitutes a lectotype designation ( ICZN 1999: Art. 74.6).

Figures

Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)

Figure 41. A Pupina dorri dorri, specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 from Haiphong, Vietnam B Pupina tongupensis, syntype NHMUK 1906.4.4.38 from Tongoop Pass, Arakan Hills, east side C Pupina anceyi, syntype MNHN-IM- 2000 - 35833 from Lang-Son and That-Khe D Pupina laffonti, syntype MNHN-IM- 2000 - 9656 from Poulo-Condor E Pupina solidula, lectotype SMF 109915 / 1 from Lang Son, Tonkin F Pupina brachysoma, syntype MNHN-IM- 2000 - 9652 from Haut Tonkin. Photo: J. Ablett, H. Taylor, NHM (B), P. Maestrati, M. Caballer, MNHN (C-F).