Arhaconotus papuanus Belokobylskij, 2000

Belokobylskij, Sergey A. & Zaldívar-Riverón, Alejandro, 2021, Reclassification of the doryctine tribe Rhaconotini (Hymenoptera, Braconidae), European Journal of Taxonomy 741, pp. 1-168 : 16-20

Arhaconotus papuanus Belokobylskij, 2000

Figs 3–4

Arhaconotus papuanus Belokobylskij, 2000: 346 .

Arhaconotus papuanus – Belokobylskij 2001: 162. — Yu et al. 2016. — Jasso-Martínez et al. 2019: 167.

Material examined

Holotype

PAPUA NEW GUINEA • ♀; “Papua New Guinea, Milne Bay Prov., Cape Vogel, Menapi, Oct. 24–29, 1989, R. Wharton ”; TAMU.

Paratypes

PAPUA NEW GUINEA • 1 ♀; same collection data as for holotype; TAMU • 1 ♂; same collection data as for holotype; ZISP .

Description

Female

MEASUREMENTS. Body length 3.7–4.3 mm; fore wing length 2.7–3.1 mm.

HEAD. Head width about 1.5 times its median length, 1.35 times width of mesoscutum. Head behind eyes (dorsal view) weakly convex in anterior half, strongly and almost linearly narrowed in posterior half. Transverse diameter of eye 2.0–2.4 times as long as temple. Ocelli medium-sized, arranged almost in equilateral triangle; POL almost equal to OD, 0.4–0.5 times OOL. Eye distinctly emarginated opposite antennal sockets, 1.2–1.3 times as high as broad. Malar space 0.3 times height of eye, 0.6–0.7 times basal width of mandible. Face width 0.8–0.9 times eye height and 1.2 times height of face and clypeus combined. Upper margin of clypeus situated distinctly higher than lower level of eyes. Width of hypoclypeal depression 0.8–1.0 times distance from depression to eye, 0.4–0.5 times width of face. Vertex convex. Head below eyes (front view) distinctly and almost linearly or weakly roundly narrowed. Antennae filiform, more then 24-segmented (apical segments missing). Scapus 1.5–1.6 times as long as its maximum width, 2.0 times length of pedicel. First flagellar segment 4.5–5.0 times as long as its apical width, 1.1 times as long as second segment.

MESOSOMA. Length about 2.0 times its height. Pronotal carina distinct, situated submedially. Mesoscutum (dorsal view) about as long as wide. Notauli deep in anterior half and shallower in posterior half, rather wide, sparsely and finely crenulate. Median lobe of mesoscutum distinctly convex anteriorly. Prescutellar depression with high median and two lateral carinae, almost smooth on rest part, distinctly curved along posterior margin, 0.3 times as long as scutellum. Scutellum length almost equal to its maximum width. Subalar depression coarsely crenulate. Sternaulus (precoxal furrow) entirely smooth. Prepectal carina with low lobes opposite fore coxae. Metanotum dorso-medially with two high and curved carinae. Propodeum weakly arcuately sloping backward (lateral view). Metapleural flange posteriorly with dense pubescence.

WINGS. Length of fore wing 3.4–3.7 times its maximum width. Pterostigma 4.0–4.5 times as long as wide. Metacarpus (1-R1) 1.1–1.2 times as long as pterostigma. Radial (marginal) cell 3.3 times as long as wide. Radial vein (r) arising from middle of pterostigma. First radial abscissa (r) forming very obtuse corner with second abscissa (3-SR). Second radial abscissa (3-SR) 3.8 times as long as first abscissa (r), 0.75–0.80 times as long as the straight third abscissa (SR1), 1.6–1.8 times as long as first radiomedial vein (2-SR). Second radiomedial (submarginal) cell long, not widened distally, 3.2–3.5 times as long as wide, about 1.7 times as long as the narrow brachial (subdiscal) cell. First medial abscissa (1-SR+M) very weakly sinuate. Second abscissa of medial vein (2-SR+M) short or very short. Recurrent vein (m-cu) 0.8 times as long as first radiomedial vein (2-SR). Mediocubital vein (M+CU1) weakly curved in apical half. Distance from nervulus (cu-a) to basal vein (1-M) 0.5–0.7 times nervulus (cu-a) length. Brachial (subdiscal) cell closed distinctly before recurrent vein (m-cu); posterior bulla present but short; posterior abscissa of anal vein (2-1A) (behind brachial vein (CU1b)) present. Hind wing 6.0 times as long as wide. Recurrent vein (m-cu) straight, strongly oblique towards base of wing.

LEGS. Hind coxa almost 2.0 times as long as wide (with tooth). Hind femur 3.2 times as long as wide. Hind tarsus 1.0–1.1 times as long as hind tibia. Hind basitarsus with lower keel and with very dense pale setosity. Second segment of hind tarsus 0.3 times as long as basitarsus, 1.1–1.2 times as long as fifth segment (without pretarsus).

MESOSOMA. Metasoma 1.4 times as long as head and mesosoma combined, with sixth visible tergites, remaining segments weakly protruding behind sixth tergite. First tergite distinctly, almost linearly and evenly widened from base to apex. Maximum width of tergite almost 2.0 times its minimum basal width; length 0.9–1.0 times its apical width, 0.8 times length of propodeum. Apical area of second tergite 0.75 times as long as remaining of tergite. Median length of second tergite (with areas) 1.0–1.2 times its basal width, almost 2.0 times median length of third tergite. Sixth tergite weakly enlarged, almost straight on posterior margin, without median emargination and postero-ventral lobes. Sixth tergite almost as long as fifth and about 1.1 times as long as fourth tergites. Ovipositor sheath about 1.1 times as long as metasoma, about 2.0 times as long as mesosoma, 0.9 times as long as fore wing.

SCULPTURE AND PUBESCENCE. Head smooth, face rugulose-striate in rather narrow vertical stripes near smooth median and lateral areas. Mesonotum and mesopleuron smooth. Metapleuron smooth and with fine reticulation or granulation in anterior half, rugulose in posterior half. Propodeum with large, mainly coriaceous, with fine rugosity along carinae and almost smooth basally basolateral areas; areola rather small, rugulose, 1.5 times as long as maximum width; median carina 2.0–2.5 times as long as anterior fork of areola; propodeum in posterior third rugulose. Hind legs smooth. First and second tergites entirely (but second in apical area finely), third tergite in basal 0.2 medially and 0.8–0.9 laterally, fourth and fifth tergites in basal furrow only medially and in basal 0.8 laterally distinctly striate. Remaining parts of tergites and sixth tergite almost entirely smooth. Second–sixth tergites laterally coarsely and densely striate but smooth apically. Vertex glabrous medially, with sparse long and semi-erect setae posteriorly and laterally. Mesonotum with dense semi-erect and rather short setae arranged wide area along notauli and marginally. Mesopleuron mostly glabrous. Hind tibia dorsally with dense numerous short setae and long sparse semi-erect setae, long setae 0.6–0.8 times as long as maximum width of tibia. Laterotergites shortly and rather sparsely setose.

COLOUR. Body black, head below and third–sixth tergites in apical halves reddish. Antenna reddish brown basally, darkened toward apex. Palps pale yellow. Legs brownish yellow, middle coxae basally, hind coxae entirely and base of hind tibia reddish brown, sometimes darker, rarely middle coxa light reddish brown; hind femur in basal half brownish. Ovipositor sheath medially light reddish brown or reddish brown, dark in basal third and apically. Fore wing distinctly and evenly infuscate. Pterostigma brown, yellow in basal basally and in apical ¼ or ⅓.

Male

MEASUREMENTS. Body length 2.9 mm, fore wing length 2.1 mm. Malar space 0.25 times height of eye and 0.55 times basal width of mandible.

WINGS. Second radial abscissa (3-SR) 5.3 times as long as first abscissa (r), 0.9 times as long as third abscissa. Second radiomedial (submarginal) cell 4.0 times as long as wide, 2 times as long as brachial (subdiscal) cell.

LEGS. Hind femur 2.7 times as long as width. Second tarsal segment of hind leg 0.25 times as long as basitarsus, 0.9 times as long as fifth segment (without pretarsus).

METASOMA. Narrow. Length of first tergite 1.35 times its apical width. Apical area of second tergite 0.6 times as long as remaining part of tergite. Length of second tergite 1.6 times its basal width, 1.8 times length of third tergite. Basal and apical areas of second tergite almost entirely smooth. Third and fourth tergites coarse striate in basal 0.6–0.7; fifth and sixth tergites reticulate-coriaceous in basal 0.7–0.8. Apical tergites not concealed by sixth tergite.

COLOUR. Body dark reddish brown. Hind femur almost entirely brownish. Otherwise similar to female.

Distribution

Papua New Guinea.

Figures

Fig. 3. Arhaconotus papuanus Belokobylskij, 2000, ♀, holotype, TAMU. A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Basal segments of antenna. E. Head, lateral view. F. Fore leg. G. Mesosoma, dorsal view. H. Mesosoma, lateral view. I. Hind coxa. J. First to third metasomal tergites, dorsal view. K. First to third metasomal tergites, lateral view.

Fig. 4. Arhaconotus papuanus Belokobylskij, 2000, ♀, holotype, TAMU. A. Wings. B. Metasoma, dorsal view. C. Metasoma, lateral view.