Geodromicus plagiatus ( Fabricius, 1798 )

Shavrin, Alexey V., 2022, The plagiatus species group of the genus Geodromicus Redtenbacher, 1857 (Insecta Coleoptera: Staphylinidae: Omaliinae), and additional data on some species from Japan, Zootaxa 5188 (5), pp. 401-429 : 404-415

Geodromicus plagiatus ( Fabricius, 1798)

( Figs. 1–7 , 12–28 )

Staphylinus plagiatus Fabricius, 1798: 180 ; Paykull 1800: 387, Fabricius 1801: 597

Lesteva plagiata: Latreille 1804: 368 , Mannerheim 1830: 56, Lacordaire 1835: 482

Anthophagus plagiatus: Gravenhorst 1802: 123, 1806: 222 ; Gyllenhal 1810: 195, Germar 1827: 5, C. Sahlberg 1827: 276, Zetterstedt 1828: 44, 1838: 47, Erichson 1840: 852, Kraatz 1857: 925

Geobius plagiatus: Heer 1839: 193

Anthophagus (Geodromicus) plagiatus: Scriba 1863: 51

Anthophagus nigritus v. plagiatus: Reitter 1870: 54

Geodromus plagiatus: Redtenbacher 1849: 747 , Fairmaire & Laboulbène 1856: 629

Geodromicus plagiatus: Redtenbacher 1857: 244, 1874: 266 , Thomson 1859: 48, 1861: 181, Fauvel 1871: 133 [= 1872: 107], J. Sahlberg 1876: 208, Mulsant & Rey 1880: 56, Ganglbauer 1895: 711, Luze 1903: 108, Sainte-Claire Deville 1907: 50, Reitter 1909: 183, Johansen 1914: 587, Porta 1926: 30, Portevin 1929: 436, Palm 1948: 121, 1961: 155, Hansen 1951: 91, Fagel 1958: 217, Smetana 1959: 358, Khnzorian 1962: 109, Lohse 1964: 62, Bordoni 1984: 24, Szujecki 2008: 132, Zanetti 2011: 67

Geodromicus (Geodromicus) plagiatus: Zanetti et al. 2008: 402

Geodromicus plagiatus v. subnigritus Lohse, 1958: 53 ; Palm 1961: 156, Zanetti 1987: 378, Herman 2001: 302 [nomen dubium]

Geodromicus pyrenaeus Jarrige, 1963: 166 ; Bordoni 1984: 27 syn. n.

Geodromicus (s.str.) lapponicus A. Bordoni, 1984: 29 [nomen dubium]

Geodromicus (s.str.) schoenmanni Bordoni, 1984: 31 ; Iablokoff-Khnzorian 1989: 137 syn. n.

Geodromicus (s.str.) klimai A. Bordoni, 1984: 32 syn. n.

Psephidonus plagiatus: Burakowski et al. 1979: 69 , Tóth 1982: 101

Psephidonus (Psephidonus) plagiatus: Zanetti 1987: 377

Lesteva globulicollis Mannerheim, 1830: 56 ; Mannerheim 1831: 470

<?!> Anthophagus verticalis Say, 1831: 49 [= 1834: 463]; Erichson 1840: 852, Fauvel 1878: 254, Casey 1893: 407, Ganglbauer 1895: 711

<?!> Lesteva canaliculatus Stephens, 1834: 360 ; Stephens 1839: 431, Waterhouse 1858: 31

Geodromicus anthracinus Brisout de Barneville, 1863: 40 ; Gemminger & Harold 1868: 658

Geodromicus plagiatus anthracinus: Kittel 1873 -1884: 168

Anthophagus globulicollis: Zetterstedt 1838: 49 , Hochhuth 1863: 102

Geodromicus globulicollis: Thomson 1861: 181 , Redtenbacher 1874: 266, J. Sahlberg 1876: 208, Ganglbauer 1895: 711 [pars], Reitter 1909: 184 [pars], Porta 1926: 30, <?!> Tottenham 1954: 36, <?!> Smetana 1959: 359, <?!> Khnzorian 1962: 110

Geodromicus plagiatus var. globulicollis: Fauvel 1871: 135 [= 1872: 109]

Other references see in Herman (2001).

Type material examined. Lectotype (here designated) of Staphylinus plagiatus Fabricius, 1798 , ♂ [specimen pinned in damaged right elytron; right antennomeres 3–11, left middle leg, right middle tarsus are missing ( Fig. 1 ); labels as in Fig. 2 ]: ’42. | plagiatus’ <handwritten>, ‘plagia- | tus’ <handwritten>, ‘TYPE’ <red, printed, pinned above narrow rectangular printed label: ‘7. 42’>, ‘ Lectotype ♂ | Staphylinus | plagiatus Fabricius, 1798 | Shavrin A.V. des. 2016’ <red, printed>, ‘ Geodromicus | plagiatus | ( Fabricius, 1798) | Shavrin A.V. det. 2016’ <printed> ( ZMUC).

Holotype of Lesteva globulicollis Mannerheim, 1830 , ♂ [dissected; left antennomeres 3–11 and right antennomeres 9–11 missing, right elytron with a hole ( Fig. 3 ); labels as in Fig. 4 ]: diamond-shaped small label without text, ‘Saimen’ <handwritten>, ‘Savon.’ <printed>, ‘Coll. Mannh.’ <printed>, ‘Mus. Hels. | N:o 3232 [handwritten]’ <printed>, ‘ Geodromicus | plagiatus F. | ab. subnigritus | Lohse | ♂ det. Thure Palm [printed]’ <handwritten>, ‘Mus. Zool. H:fors [printed] | Spec. typ. No [printed] 326. | Anthophagus glo= | bulicollis Mann.’ <handwritten>, ‘http:// id.luomus.fi/ | GAC.20090 | FINLAND Sa Saimaa | 61.30 N, 28,19 E | Mannerheim leg.’ <printed, with barcode on left side of the label>, ‘Photographed| 2016 | Pekka Malinen’ <blue, printed>, ‘ Geodromicus | plagiatus | ( Fabricius, 1798) | Shavrin A.V. det. 2022’ <printed> ( MZHF).

Holotype of Geodromicus plagiatus a. subnigritus Lohse, 1958 , ♂ [dissected]: ‘Taurach bei | 15.IX.[19]58 [handwritten] | Lintsching Lungau’ <rectangular label, printed in blue>, ‘Typus’ <red rectangular label, handwritten by blue pen>, ‘ Geodromicus | plagiatus | v. subnigritulus [sic!]’ <handwritten by blue pen, with black margin>, ‘Coll. | G.A. Lohse | MHNG-1994’ <printed>, ‘ Geodromicus | plagiatus ( Fabricius, 1798) | Shavrin A.V. det. 2015’ <printed> ( MHNG). Paratypes: 1 ♂ [dissected], 1 ♀: ‘Spittal [an der] Drau’ <handwritten by red pen>, ‘paratypoid’ <red, handwritten by blue pen>, ‘Coll. | G.A. Lohse | MHNG-1994’ <printed>, ‘ Geodromicus | plagiatus ( Fabricius, 1798) | Shavrin A.V. det. 2015’ ( MHNG).

Holotype of Geodromicus pyrenaeus Jarrige, 1963 , ♂ [dissected]: ‘p.2’ <handwritten>, ‘Holçarte | B. P. IX.[19]55’ <handwritten>, ‘TYPE’ <red, printed>, ‘ Geodromicus | pyrenaeaus Jarrige, 1963 | Shavrin A.V. det. 2017’ <printed> ( MHNH). Paratypes: 1 ♂, 1 ♀: ‘Larrau | Basses Pyrénées | 25. VI. 1955 | L. Levasseur’ <handwritten>, ‘ Geodromicus | pyrenaeus Jarr. | (paratype)’ <handwritten>, ‘Para- | type’ <red label, with thin black frame>, ‘ Geodromicus | plagiatus ( Fabricius, 1798) | Shavrin A.V. det. 2018’ <printed> ( IRSNB).

Holotype of Geodromicus (s.str.) lapponicus Bordoni, 1984 , ♂ [dissected; specimen damaged: left antenna, left tarsus and tibia of fore leg, right middle and hind left legs missing]: ‘Laponie’ <handwritten>, ‘Coll. et det. A. Fauvel | Geodromicus [handwritten] | globulicollis Mannh. [handwritten] | R I. Sc.N.B. 17.479’ <printed>, ‘Syntype’ <printed>, ‘Holotypus’ <red, printed>, ‘ Geodromicus | lapponicus n.sp. [handwritten] | Det. A. Bordoni 1983 | holot. ♂ ’ <printed> ( IRSNB).

Holotype of Geodromicus (s.str.) schoenmanni Bordoni, 1984 , ♂ [dissected]: ‘ ♂ ’ <printed>, ‘Kaukasus’ <handwritten>, ‘ex coll. | Luze’ <yellow, printed>, ‘HOLOTYPUS’ <red, printed label with black margin>, ‘ Geodromicus | schoenmanni n. sp. | Det. A. Bordoni 19[printed]83 [handwritten above dots] | holotypus’ <handwritten>, ‘ Geodromicus | plagiatus ( Fabricius, 1798) | Shavrin A.V. det. 2015’ <printed> ( NMW). Paratypes: 1 ♂ [left antennomeres 5–11, right antennomeres 2–11 missing, apical part of elytra damaged; apical abdominal tergite VIII and sternite VIII and male genital segment glued on the same card under the beetle; dissected], 1 ♂ [right antennomeres 3–11, right middle leg and aedeagus are missing; dissected]: ‘ ♂ ’ <printed>, ‘Kaukasus’ <handwritten>, ‘ex coll. | Luze’ <yellow, printed>, ‘PARATYPUS’ <red rectangular printed label, in thin black frame>, ‘ Geodromicus | schoenmanni | n. sp. | Det. A. Bordoni 19[printed]83 | Holotypus’ <handwritten> ( NMW); 1 ♀: ‘ ♀ ’ <printed>, ‘Kaukasus’ <handwritten>, ‘ex coll. | Luze’ <yellow, printed>, ‘PARATYPUS’ <red label, with black margins, printed>, ‘ Geodromicus | schoenmanni | n. sp. | Det. A. Bordoni 19[printed]83 [handwritten above dots] | Holotypus’ <handwritten> ( NMW). All paratypes with additional label: ‘ Geodromicus | plagiatus ( Fabricius, 1798) | Shavrin A.V. det. 2015’.

Holotype of Geodromicus (s.str.) klimai A. Bordoni,1984 , ♂ [dissected]:‘ ♂ ’<printed>,‘Kaukasus’<handwritten>, ‘ex coll. | Klima’ <blue, printed>, ‘ Geodromicus | schoenmanni n. sp. | Det. A. Bordoni 19[printed]83 [handwritten above dots] | holot.’ <handwritten>, ‘HOLOTYPUS’ <red label with black margins, printed> ( NMW).

Additional material examined. NORWAY: FINNMARK: 1 ♂: Lakselv. 20.06- 04.07.2012. N. Savenkov leg. ( DUBC); 1 ♂: Lakselv. 29.07.1978. G. Gillerfors leg. ( ZML); 1 ♂, 1 ♀: Vassbyggda i Aurlandsdal. 100 m a.s.l. 15.08.1965. C.H. Lindroth leg. ( ZML); 3 ♀♀: Bossekop. 07.1898 ( BMNH); 1 ♀: Adamsfossen. 30.07.1978. G. Gillerfors leg. ( ZML); 1 ♀: Karasjoki. U. Sahlberg leg. ( MZHF); 1 ♂, 2 ♀♀: Karasjok. 1907. T. Münster leg. ( MZHF); TROMS: 1 ♂: Rundhaug. 27.06.1930. F. Jensen leg. ( ZMUN); 1 ♂: ‘Moen 3/783’ ( MZHF); 1 ♀: Nordreisa Norge. 12- 28.06.1954. O. Sjöberg leg. ( ZML); 1 ♂: Målselva, Rundhaug. Klefbeck leg. ( ZML); 2 ♀♀: Aronnes. 06- 07.08.1960. G. Israelson leg. ( ZML); NORDLAND: 1 ♂: Vestfjord dal. T. Münster leg. ( ZMUN); 1 ♂: Erlkstadfj, Lødingen. T. Münster leg. ( ZMUN); 1 ♂: Lødingen. T. Münster leg. ( ZMUN); 1 ♂: Vestfjord. T. Münster leg. ( ZMUN); SØR-TRØNDELAG: 1 ♀: Ålen ( IRSNB); 5 ♂♂: Vassbygda. 15.08.1965. I. Svensson leg. ( ZML); OPPLAND: 1 ♀: Norse Skjak. < Geodromicus globulicollis > (cR); 1 ♂: Skjak, Marlobrua. 27.06.1978.A. Fjellberg leg. ( ZMUN); 1 ♂, 1 ♀: Vågåmo. 1953. T.E. Leiler leg. ( SMNH); 1 ♂: Gjøvik. 31.07.1896. Warloe leg. ( ZMUN); 1 ♂: same data, 17.07.1897 ( ZMUN); 2 ♂♂, 1 ♀: Fagernaes. T. Münster leg. ( FMNH); 1 ♂: Gjøvik. 31.07.1896. Warloe ( ZMUN); BUSKERUD: 5 ♂♂: Sørum, Våge. 06.1925. T. Münster leg. ( ZMUN); HORDALAND: 4 ♂♂: Røldal ( ZMUN); OSLO: 1 ♂: Sokna, Støren. 20.07.1917. T. Münster leg. ( ZMUN); TELEMARK: 1 ♂: Fyresdal. 05.1919. T. Münster leg. ( ZMUN); ROGALAND: 1 ♀: Arnoy. 06-07.1958. P.J.M. Greenslade leg. ( BMNH); 1 ♂: Klinken. 16.08.1969. R. Baranovski ( ZML); REGION NOT LOCATED: 1 ♂: ‘Norwege Schuster’ ( MHNH); 1 ♂: ‘Norwege’ ( MHNH); SWEDEN: LAPPLAND: 4 ♂♂: Abisko. 07.1957 ( MHNG); 1 ♀: Reivo. 23.08.1989. G. Gillerfors leg. ( ZML); 1 ♀: Njuonjes, Kvikkj-Njuonjesberget. 22.07.1924. E. Arvidson leg. ( ZML); 1 ♂: Kalnisjokk bei Messaure, 66°42‘N 20°25‘E. 18.7.1967. E. Thomas leg. ( MHNG); JäMTLAND: 1 ♂: Vilhelmina. 21.07.1936. C.H. Lindroth leg. ( ZML); 1 ♂: Långflon. 15.06.1933. C.H. Lindroth leg. ( ZML); 1 ♀: Bispgården. 03.06.1930. C.H. Lindroth leg. < plagiatus nigrita > ( ZML); MEDELPAD: 1 ♂: Lidensboda. 06- 11.07.1937. Bruce leg. ( ZML); VäSTERBOTTEN: 1 ♂: Luleå, Karlsvik. 18.06.1938. C.H. Lindroth leg. ( ZML); 1 ♀: Vistbäcken. 02.07.1979. G. Gillerfors leg. ( ZML); DALARNA: 1 ♂: Kringetpjarden. 23.06.1974. G. Gillerfors leg. ( ZML); 2 ♂♂: Särnaheden. 14.08.1969. R. Baranovski leg. ( ZML); 1 ♂: Sardaes. 06.1920. T. Münster leg. ( ZMUN); HäLSINGLAND: 1 ♀: Ramsjö. 17.06.1971. R. Baranovski leg. ( ZML); VäRMLAND: 1 ♀: Degerfors. 22.07.1957. Welden leg. ( ZML); 1 ♀: Höljes. 07.08.1981. R. Baranovski leg. < nigrita > ( ZML); 1 ♀: Vrml. Åstrand. 11.08.1968. R. Baranovski leg. ( ZML); UPPLAND: 1 ♂: Grisslehamn Karlsand. 12.09.1969. R. Baranovski leg. ( ZML); 1 ♀: Marskär. 10.06.1959. G. Gillerfors leg. ( ZML); SÖDERMANLAND: 1 ♂: Tuna. 10.07.1997. T. Kvamme leg. ( ZMUN); ÖSTERGÖTLAND: 1 ♂: Tåkern ( ZMUC); 1 ♂: Lysingshdsallm. T. Palm leg. ( ZMUC); 1 ♀: ‘Jotunfj[ord].’, ‘Suecia’ ( SNMB); 2 ♂♂, 1 ♀: ‘Omb. tr. Vättern’ ( ZML, ZMUC); 1 ♂: Lönsås. 30.08.1938. S. Gaunitz leg. ( ZML); VäSTER-GÖTALAND: 1 ♀: Öglunda. 25.06.1943. P. Brinck leg. ( ZML); 2 ♀♀: Djupvik. 19.07.1969. R. Baranovski leg. ( ZML); GOTLAND: 1♂: Wisby, seaside. 03.08.2013. M. Balalaikins leg. ( DUBC); REGION NOT LOCATED: 1 ex.: ‘Suecia’ ( ZIN); FINLAND: LAPLAND: 12 ♀♀: Utsjoki, Tenjoki. U. Saalas leg. ( MZHF); 3 ♂♂, 1 ♀: same locality. U. Sahlberg leg. ( MZHF); 1 ♂, 1 ♀: same locality. 23.06.1973. O. Biström leg. ( MZHF); 1 ♀: Tenojoki. U. Sahlberg leg. ( MZHF); 2 ♂♂: Inari. B. Poppius leg. ( MZHF); 3 ♂♂, 1 ♀: same locality. O. Renkonen leg. ( MZHF); 1 ♀: same locality. S. Platonoff leg. ( MZHF); 1 ♀: same locality. R. Krogerus leg. ( MZHF); 1 ♀: same locality. E. Helve leg. ( MZHF); 6 ♂♂, 6 ♀♀: Enare. O. Biström leg. ( MZHF); 3 ♂♂, 3 ♀♀: Enontekiö. O. Renkonen leg. ( MZHF); 1 ♂: same locality. J. Sahlberg leg. ( MZHF); 1 ♀: same locality. U. Saalas leg. ( MZHF); 1 ♂: Pallastunt. U. Sahlberg leg. ( MZHF); 1 ♀: Pallasjärvi. 14.07.1957. A. Wegelius leg. ( MZHF); 1 ♀: Inari, Tenjoki. U. Saalas leg. ( MZHF); 1 ♂, 2 ♀♀: Inari. E. Helve leg. ( MZHF); 1 ♂: Inarjoki. U. Sahlberg leg. ( MZHF); 2 ♂♂: Kusräka ( FMNH); 3 ♀♀: Lutto. S. Platonoff leg. ( MZHF); 6 ♂♂: Salla. 24.06.1937. E. Kangas leg. ( MZHF); 2 ♂♂, 2 ♀♀: same locality. S. Stockmann leg. ( MZHF); 3 ♀♀: Palojoki. J. Sahlberg leg. ( MZHF); 1 ♀: Saariselkä. B. Poppius leg. ( MZHF); 1 ♂, 1 ♀: Muonio. F.G. Mäklin leg. ( MZHF); 1 ♀: same locality. V. Alava leg. ( MZHF); 2 ♀♀: same locality. M. Hellén leg. ( MZHF); 1 ♀: Kittilä. U. Sahlberg leg. ( MZHF); 1 ♂: Enontekiö. U. Sahlberg leg. ( MZHF); 5 ♂♂: Rovaniemi. 25.7.1956. H. Lindberg leg. ( MZHF); 1 ♂, 1 ♀: Kemijärvi Pyhäjoki. 30.06.1951. H. Lindberg leg. ( MZHF); OULU: 22 exs.: Petsamo. 02- 23.07.1929. H. Lindberg leg. ( MZHF); 1 ♂, 1 ♀: same locality. E. Thuneberg leg. ( MZHF); 2 ♂♂: Oulu. Y. Wuorentaus leg. ( MZHF); 1 ♂: Uleåborg. J. Sahlberg leg. ( MZHF); 1 ♀: Sotkamo. J. Carpelan leg. ( MHNG); 1 ♂: Kalajoki ( MZHF); 2 ♂♂, 1 ♀: Säräsniemi. Y. Wuorentaus leg. ( MZHF); 4 ♂♂, 5 ♀♀: Kajaari. J. Carpelan leg. ( MZHF); 2 ♂♂, 1 ♀: Kuusamo. S. Stockmann leg. ( MZHF); EASTERN FINLAND: 1 ♂: Pielisjärvi. 31.07.1970. R. Baranovski leg. ( ZML); 1 ♀: same data. 01.08.1970 ( ZML); 32 exs.: Keitele, 62º53`N 25º58`E. 08.1997. P.M. Hammond leg. ( BMNH); 1 ♀: Maaninka. 27.07.1940. R. Elfving leg. ( MZHF); 1 ♂, 1 ♀: Salmi. S. Platonoff leg. ( MZHF); 1 ♀: Metsäpirtti. M. Hellén leg. ( MZHF); 1 ♂: same locality. P.H. Lindberg leg. ( MZHF); 1 ♀: same locality. J. Sahlberg leg. ( MZHF); 1 ♂, 1 ♀: same locality. M. Häyrynen leg. ( MZHF); 2 ♂♂: Tyrvää. 26.07.1972. P. Savolainen leg. ( MZHF); 1 ♂: Asikkala. 08.1937. Hellman leg. ( MZHF); 1 ♂: same locality, Tuoomikoski leg. ( MZHF); 1 ♂: Ristiina. Linnaniemi leg. ( MZHF); 9 ♂♂, 6 ♀♀: same locality. O. Renkonen leg. ( MHNG, MZHF, SNMB); 1 ♂: Velmersalmi. Hemdal leg. ( MZHF); 1 ♂: Maaninga. Lundström leg. ( MZHF); 1 ♂: Kuopio. Wellenius leg. ( MZHF); 1 ♂, 1 ♀: same locality. K.M. Levander leg. ( MZHF); 1 ♂: Nilsiä. K.M. Levander leg. ( MZHF); 1 ♀: Idensaimi. J. Sahlberg leg. ( MZHF); 1 ♂, 2 ♀♀: Vermersalmi. Hemdal leg. ( MZHF); 2 ♂♂, 1 ♀: Kontiolahti. 19.08.1942. H. Lindberg leg. ( MZHF); 2 ♂♂: same data. 24.08.1942 ( MZHF); 2 ♂♂, 1♀: same data, 29.08.1942 ( MZHF); 1 ♂: Eno. M. Laamanen leg. ( MZHF); 1 ♂, 1 ♀: Kitee. 27- 29.06.1963. P. Savolainen leg. ( MZHF); 1 ♀: Karislojo. J. Sahlberg leg. ( MZHF); 2 ♂♂, 1 ♀, 17 exs.: Kirjavala. B. Poppius leg. ( MZHF); 2 ♂♂: Parikkala. 14- 15.06.1956. H. Lindberg leg. ( MZHF); WESTERN FINLAND: 1 ♂, 1 ♀: Yläne ( IRSNB); 3 ♂♂, 7 ♀♀: same locality. U. Saalas leg. ( MZHF); 1 ♀: same locality. J. Sahlberg leg. ( MZHF); 3 ♂♂, 6 ♀♀: Karstula. M. Pohjola leg. ( MZHF); 7 ♂♂, 3 ♀♀: Kangasala. A. Saarinen leg. ( MZHF); 1 ♀: Uusikaup. Söderman leg. ( MZHF); 1 ♀: Parkano. 18.08.1928. R. Elfving leg. ( MZHF); 1 ♀: Ytterö. 22.6.1934. R. Elfving leg. ( MZHF); 1 ♂: Pikkala. Grönblom leg. ( MZHF); 1 ♂, 2 ♀♀: Itä-Teisko. A. Saarinen leg. ( MZHF); 1 ♂: Lempäälä. Hemdal leg. ( MZHF); 2 ♂♂, 1 ♀: Hattula. A. Wegelius leg. ( MZHF); 11 ♂♂, 6 ♀♀: same data. 01.09.1947 ( MZHF); 1 ♀: Pielavesi. E. Palmén leg. ( MZHF); 1 ♂: Konginkangas. Luther leg. ( MZHF); 1 ex.: Luopioinen. 23.08.1952. E. Kangas leg. ( MZHF); 1 ♂: Ruovesi. J. Sahlberg leg. ( MZHF); 1 ♂, 2 ♀♀: Karkku. M. Hellén leg. ( MZHF); 1 ♂: Pori. 24.06.1962. L. Tiensuu leg. ( MZHF); 1 ♂: Vammala. 20.07.1962. P. Savolainen leg. ( MZHF); 3 ♂♂, 1 ♀: Pirkkala. O. Wellenius leg. ( MZHF); 1 ex.: Virrat. E. Kangas leg. ( MZHF); 1 ♀: Pihlajavesi. J. Sahlberg leg. ( MZHF); 1 ♂: Nurmiaho. J. Sahlberg leg. ( MZHF); SOUTHERN FINLAND: 1 ♂: Eckerö. Forsius leg. ( MZHF); 1 ♂, 3 ♀♀: same locality, H. Lindberg leg. ( MZHF); 1 ♂, 1 ♀: Kökar. 07.08.1941. H. Lindberg leg. ( MZHF); 2 ♂♂: same locality. S. Stockmann leg. ( MZHF); 4 ♀♀: Sottunga Husö. 3.7.1942. H. Lindberg leg. ( MZHF); 1 ♀: same data. 08.1943 ( MZHF); 2 ♂♂: Jomala. S. Stockmann leg. ( MZHF); 2 ♂♂, 1 ♀: same locality. S. Platonoff leg. ( MZHF); 1 ♂: Lemland. S. Stockmann leg. ( MZHF); 4 ♂♂, 2 ♀♀: Karislojo. J. Sahlberg leg. ( MZHF); 1 ♂: Karjalohja. 01.07.1938. O. Renkonen leg. ( MZHF); 1 ♂, 1 ♀: Lohja. Linnaniemi ( MZHF); 1 ♀: Birkkala. O. Wellenius leg. ( MZHF); 1 ♀: Hörikkä. Leikko leg. ( MZHF); 1 ♀: Kustö. Holmquist leg. ( MZHF); 1 ♂: Tvärminne. U. Saalas leg. ( MZHF); 1 ♂, 1 ♀: Lojo. 13.07.1917. H. Lindberg leg. ( MZHF); 4 ♂♂, 4 ♀♀: same data. 21.06.1916 ( MZHF); 1 ♀: Sammatti. J. Sahlberg leg. ( MZHF); 6 ♂♂, 1 ♀: Tvarminne. Axelson leg. ( MZHF); 1 ♂: same locality. Y. Wuorentaus leg. ( MZHF); 1 ♂: same locality. H. Lindberg leg. ( MZHF); 2 ♂♂: same locality, 23.6.1975. O. Biström leg. ( MZHF); 1 ♀: same data. 26.08.1975 ( MZHF); 1 ♀: same data. 30.08.1976 ( MZHF); 1 ♀: same data. 27.9.1975 ( MZHF); 1 ♀: same data. 30.05.1975 ( MZHF); 1 ♀: same data. 12.6.1975 ( MZHF); 1 ♂, 1 ♀: Espoo. E. Helve leg. ( MZHF); 1 ♂, 1 ♀: Sibbo. S. Stockmann leg. ( MZHF); 1 ♀: Pasila. Winter leg. ( MZHF); 1 ♂: Valkjärvi. 14.8.1934. P.H. Lindberg leg. ( MZHF); 1 ♂: Esbo. O. Biström leg. ( MZHF); 2 ♀♀: Muolaa. E. Palmén leg. ( MZHF); 1 ♀: Helsingfors. A.G. Helenius leg. ( MZHF); 1 ♀: Borgå Onas. S. Stockmann leg. ( MZHF); 1 ♂, 3 ♀♀: Luumäki. V. Numers leg. ( MZHF); 1 ♀: Joutseno. 12- 13.06.1956. H. Lindberg leg. ( MZHF); 1 ♀: same locality. E. Thuneberg leg. ( MZHF); 1 ♀: Virolakti. 03.06.1975. T. Clayhills leg. ( MZHF); 1 ♂, 1 ♀: Sakkola. M. Hellén leg. ( MZHF); REGION NOT LOCATED: 1 ♀ ‘Finlande’ ( IRSNB); 2 ♀♀: ‘Lapponia’, ‘ G. globulicollis’ ( MZHF); BELGIUM: 3 ♀♀: Hockai 22.07.1966. R. Van Dorsselaer leg. ( MHNG); FRANCE: HAUTS-DE-FRANCE: 1 ♀: Ecottes ( IRSNB); AUVERGNE-RHȎNE-ALPES: 1 ♂: Le Lioran. 01.08.1926. ( MHNH); 1 ♂, 1 ♀: same locality. 28.08.1912. G. Tempère leg. ( MHNG); 1 ♀: same locality. 06.1929 ( MHNH); 2 ♂♂: same locality. 12.07.1939. E. Alraud leg. ( MHNG); 1 ♂: same locality. 1200 m a.s.l. 15.06.1946. L. Levasseur leg. ( MHNH); 1 ♂, 4 ♀♀: Vallorcine. 07.1959. ( MHNH); 1 ♀: Averole. 2000 m. ( MHNH); 1 ex.: Salève. 08.09 ( ZIN); 1 ♂: Torrent du Guiers route du Sapey < Geodromicus plagiatus v. nigrita > ( IRSNB); 1 ♀: Briancou. 12.06.1965. A. Comellini leg. ( MHNG); 3 ♂♂, 1 ♀: Valloires. 27.07.1939. L. Giraud leg. ( MHNG); 2 ♂♂: Thonon. 09.1959 ( MHNG); 4 exs.: Villard d`Arene. 1700 m. 08.1954. ( MHNH); 1 ♂: La Bérarde. 1700 m. 08.1963. J. Jarrige leg. ( MHNH); 1 ♀: Ailefroide. 1700 m a.s.l. 03.08.1947. R. Lavallard leg. ( MHNH); 1 ♀: Col De Vars. 07.1962. ( MHNH); 1♂: Lanans. 09.1955 ( MHNH); OCCITANIA: 1 ♂, 1 ♀: Lac Bleu. 06.1945. Ch. Fagniez leg. ( MHNH); 2 ♀♀, 2 exs.: Vernet. 600 m a.s.l. 06.1958. L. Levasseur leg. ( MHNH); 3 ♂♂, 3 ♀♀: Layolle. 17.07.1957 ( MHNG); 1 ♂: Canigou. 600 m a.s.l. 05.1956 ( MHNH); 1 ♀: Canigou P.O. Marialle, mountes de Clipodere. 1800 m a.s.l. 06.1952 ( MHNH); NOUVELLE-AQUITAINE: 1 ♂: Gavarnie Cascade. 13.08.1953 ( MHNG); 2 ♀♀: Gavarnie. 1400 m a.s.l. 07.1937. G. Tempère leg. ( MHNG); 1 ♂, 2 ♀♀: Gavarnie ( IRSNB); 1 ♂: Ariege, Erce Bords du Genhet. 05.09.1961. H. Coiffait leg. ( MHNH); 1 ♀: Lasqoux Vallua d`Orde. 05.08.1954. H. Coiffait leg. ( MHNH); 1 ♂: Encé. 05.01.1967. H. Coiffait leg. ( MHNH); 1 ♂, 20 exs.: Larrau. 25.06.1955. L. Levasseur leg. ( MHNH); 1 ♂: same data. 25.06.1955 ( IRSNB); 3 ♂♂, 3 ♀♀: Holçarte. 09.1955 ( MHNH); 1 ♂: same data, 07.1967 ( MHNH); 1 ♂: Bious Artiguer. 1400 m a.s.l. 11.07.1969 < Geodromicus pyrenaeus M. Tronquet det.> (cSh); SPAIN: CATALUñA: 1 ♂, 1 ♀: Cerdagne ( FMNH); 1 ♂, 2 ♀♀: Lio, haute vallée du Sègre. 02.08.1971. G. Tempère leg. ( MHNG); 1 ♀: Puerto de Roses. 1500-1800 m a.s.l. 14.05.1981. Zwick leg. ( MHNG); SWITZERLAND: BASEL: 1 ♀: Basel ( SF); BERN: 1 ♀: Le Forges bei Undervelier. 15.08.1977. V. Puthz leg. ( MHNG); 1 ♀: Habkern, Lombachzufluss. 1340 m a.s.l. 03.08.1979. V. Puthz leg. ( MHNG); VAUD: 1 ♀: ‘Pic Chaussy Südschweiz Dr. K. Singer’ ( FMNH); VALAIS: 1 ♂, 12 ♀♀: Vals. 07.08.1922. ( NHMB); 1 ♀: Vals, Raetia. 1932. Joerger leg. ( NHMB); 1 ♂: Zinal ( BMNH); 2 ♂♂: Hinterrhein. 2050 m a.s.l. 14.07.1953. O. Lundblad leg. ( SMNH); 1 ♀: ‘Vals, Raetia Joerger. [19]32’ ( NHMB); 1 ♂: Jaas Valais ( MHNH); GRAUBüNDEN: 1 ♂: Davoser Berge. 19.08.1934. Wolf leg. ( MHNG); 2 ♂♂: Vals Grison ( MHNH); 1 ♀: Pontresina ( MHNH); 1 ♂, 1 ♀:Arosa. 07.1899 < v. nigrita Müll. det. Bernh.> ( FMNH); GERMANY: SCHLESWIG-HOLSTEIN: 1 ♂: Klinken. 16.08.1969. R. Baranovski leg. ( ZML); NIEDERSACHSEN: 1 ♀: Oderbrück Horz Ufer der Oder. 04.07.1970 ( MHNG); THURINGIA: 1 ♀: Watburg. 27.06.1976. G. Gillerfors leg. ( ZML); BADEN-WüRTTEMBERG: 3 ♂♂, 3 ♀♀: ‘Ulm Württbg.’ ( ZMM); 1 ♂: Karidol. 1934. Winagrith leg. ( FMNH); BAVARIA: 1 ♂: München. 28.05.1986 ( MHNH); 1 ♂: Krün. 24.06.1948. E. Weise ( MHNH); 2 ♂♂: Erzgebinge < v. nigrita > ( FMNH); 3 ♂♂: Erlangen ( IRSNB); 1 ♂, 1 ♀: Oberau ( MHNG); 1 ♂, 1 ♀: Regensburg. 28.09.1911. Benick leg. ( MHNG); 1 ♂: Königsee. 13.06.1907 ( MHNG); 1 ♀: Baviere ( IRSNB); REGION NOT LOCATED: 3 ♂♂, 1 ♀: ‘Allemagne’ ( IRSNB); 3 ♀♀: ‘Saxon’, ‘Boheman’ ( SMNH); CZECH REPUBLIC: LIBEREC REGION: 1 ♂: ‘Nový Svĕt, VII-VIII. Krkonoše Boh[emia]. b[oreal]. Dr. Obenberger lgt.’ ( NMPC); MORAVIA-SILESIA: 1 ♂, 1 ♀: Moravia, Ostrava. S. Jureček leg. ( NMPC); 1 ♂, 1 ♀: Bezkydy. S. Jureček leg. ( NMPC); 2 exs.: Lysa Hora. 07.1903. Wagner leg. ( ZIN); 1 ♂: ‘ plagiatus Moravia Schreiber’ ( FMNH); REGION NOT LOCATED: 1 ex.: ‘Bohemia Reitter’ ( ZIN); 1 ♂: ‘Bohemia’, ‘Staudinger’ ( ZMM); 7 exs.: ‘Moravia Reitter’ ( MHNH, ZIN); SLOVAKIA: TRENčIN: 3 ♀♀: ‘C. Trencsén [Trenčin] Dr. Brancsik’ ( FMNH); PREŠOV: 4 exs.: ‘Dunajec’, ‘Slow[enian]. Pieniny’, ‘Roubal’ ( SNMB); 1 ♀: Lubotin. 1959. A. Smetana leg. ( BMNH); REGION NOT LOCATED: 1 ♀: ‘G. Strauss Nied[ere]. Tatra’ ( FMNH); 2 ♂♂: ‘Slovakia Pieniny’ ( MHNG); AUSTRIA: TYROL: 1 ♂: Lechtal Schwarzwasserbach Mündung Umgebung. 900 m a.s.l. 26.05.1995 C.M. Brandstetter leg. (cZan); 1 ♂: St. Martin. 31.07.1998. Eifler leg. (cA); 1 ex.: ‘Tauters Tir[ol]. Luze’ ( ZIN); 1 ex.: ‘Steyr O.-D. Juni 1940 Bernhauer’ ( FMNH); 1 ♂: Südtirol, Aner. 16.07.1961 ( MHNG); 1 ♂: Sölden, Ötztal 26.07.1973 ( MHNG); 1 ♂, 1 ♀: ‘Tirol B[e]r[n]h[auer] Innichen’ ( FMNH); 1 ♀: Pillerbach. 04.08.1967. Th. Palm leg. ( ZML); 6 ♂♂, 1 ♀: Götzens. 1500-2000 m a.s.l. 24.06.- 06.07.1969. Th. Palm leg. ( ZML); 1 ♂: Lechtal Schwarzwasserbach Mündung Umgebung. 900 m a.s.l. 26.05.1995. C.M. Brandstetter leg. (cZan); 1 ♂: St. Martin. 31.07.1998. Eifler leg. (cA); 1 ♂: Pusterthal < v. nigrita Müll. det. Bernh.> ( FMNH); 1 ♂: Aner. 16.07.1961 ( MHNG); 1 ♀: Holern bei Auer. 23.07.1973 ( MHNG); 1 ♂: Stoder < v. nigrita det. Bernh.> ( FMNH); 1 ♂: Nassfeld. 1600 m a.s.l. M. Bernhauer ( FMNH); OBERÖSTERREICH: 2 ♂♂: ‘Gr. Hollenstein N.-Oe. Bernh.’, ‘Ufer’, ‘August’ ( FMNH); 1 ♀: ‘Hollenstein Lewisel’ < v. nigrita Mül. > ( FMNH); 1 ♀: ‘Grünburg, O. Oe. Bernhauer’ < plagiatus det. Bernh.> ( FMNH); SALZBURG: 1 ♂: Ostermiething. Leder leg. ( MHNG); 1 ♀: ‘Hohe Tauern Koetschachtal Bernhauer’ ( FMNH); 4 ♂♂, 8 ♀♀: ‘Badgastein Gravenegg Bernhauer’ ( FMNH); 1 ♂: ‘Anlauftal Aachenfer 1300 m | II. [19]33’ ( FMNH); 1 ♂: ‘Grünb[ur]g. O[estlich].Oe[sterreich]. Sand. Bernh.’ ( FMNH); 1 ex.: ‘Gasteinertal Bad Bruck 850 m Bernh.’ ( FMNH); 1 ex.: ‘Gastein-Rtal Bad Bruck 850 m Bernh[auer].’ ( FMNH); KäRNTEN: 1 ♂: Nauthen im Gailtal. 19.07.1973. ( MHNG); 11 ♂♂, 6 ♀♀: same data. 21.7.1973. ( MHNG); 1 ♂: Gailtal, Karut. 06.1929 ( FMNH); 1 ♂, 2 ♀♀: Gailtal Cr. K. Klimsch ( MZHF); 1 ♂: ‘Carinthia, Savin. F. Ramboušek’ ( NMPC); 1 ♂: ‘Carinthia Sachsenburg’ ( NMPC); 1 ♂: Sölden. Ötztal. 26.07.1973 ( MHNG); 1 ♀: Wochein.Moczarski < v. nigrita det. Bernh.> ( FMNH); 1♂: Mallnitz. 20.06.1930.Köller leg. ( MHNG); STEIERMARK: 1 ♀: ‘Steiermark’ ( FMNH); VIENNA: 2 exs.: ‘Umgeb. von Wien’ ( ZIN); 1 ♂: ‘env. Vienna’ ( MHNG); 1 ♂: ‘Autrice env. Wien’ ( MHNG); ITALY: PIEMONTE: 1 ex.: Meeralpen, Terme de Valdieri. 11.07.1974 ( MHNG); VENETO: 1 ♂: Piave River, 46°0.58’N 11°59.9’E. 18.08.1997. J. Weipert leg. ( NME); LOMBARDIA: 2 ♂♂, 1 ♀: Turbigo [Ticino River] Ca di Bis. 15.05.1921 (cZan); 1 ♂: Ticino Pavia. 27.09.1919 ( MHNG); TRENTO: 1 ♂: Smarano, Loc. Vordes. 1000 m a.s.l. 23.07.1969. A. Zanetti leg. ( MHNG); UMBRIA: 1 ♀: Cascia dint Umbria. 09.1961. Battoni leg. ( MHNG); SLOVENIA: 2 ♀♀: ‘Wochein [Bohinj], Car[niolia]. Moczarski’ ( FMNH); 1 ♂, 1 ♀: ‘Wochein [Bohinj] Carniolia Bernhauer’ ( FMNH); HUNGARY: ‘Hungaria bor. Marmaros Reitter’ ( MHNH); POLAND: ŚWIęTOKRZYSKIE VOIVODESHIP: 2 ♂♂, 2 ♀♀: ‘Netolitzky […illegible…]’, ‘Kreis Kielce J. Netolitzky’, ‘Bogorya’ ( SDEI); WOJEWÓDZTWO MAłOPOLSKIE: 1 ♀: ‘Polen. 7.VII.[19]27 Rabka [Rabka-Zdrój] lg. Tenenbaum’ ( FMNH); 1 ♀: ‘Tatra Zakopane’ ( ZMM); WOJEWÓDZTWO DOLNOŚLąSKIE: 1 ♀: ‘Silesia Teschen Th.V. Wanka’ ( MHNG); ROMANIA: CARAş-SEVERIN: 1 ♂:‘Banatanina’ ( FMNH); SIBIU: 1 ♀: 2 km S Cǎrţişoara, bank of stream Laiţa, near bridge, 1 km W stream Cǎrţişoara. 45º42’17’’N 24º34’11’’ E. 520 m a.s.l., flotation from sand/gravel and debris on bank. 16.05.2003 Gy. Makranczy leg. (cZan); CLUJ: 1 ♂: Munţii Vlădeasa, Rogojel, 5 km SW Sãcuieu, bend of p. Alunis under hill, 46º47’38’’N 22º49’59’’E. 920 m a.s.l, plant debris stuck at branches in stream, flotation [300]. 9.06.2007. Gy. Makranczy leg. ( HNHM); HARGHITA: 1 ♂: Munţii Hăsmaş, Cheile Bicazului, 1 km E Pt. Altarului 3 km ESE Bicaz-Chei, 46º48’45’’N, 25º49’25’’E. 900 m a.s.l., from fine gravel at fast rocky stream, flotation [122]. 27.06.2004. Gy. Makranczy leg. ( HNHM); ALBA: 7 ♂♂, 8 ♀♀: Munţii Bihor, Gĭrda (de Sus), confluence of p. Dizditu and Arieşu Mare, 46°27’25’’N 22°51’02’’E. 720 m a.s.l., from mosses on stones at streambank, flotation [222]. 30.06.2006. Gy. Makranczy leg. ( HNHM); HUNEDOARA: 1 ♂: P.N. Retezat, 2 km W Lunca Rotunda, Lăpuşnicu Mare, 45°19.0’N 22°45.5’E. 1100 m a.s.l., ex streambank sand & fine gravel, floation. 29.05.2002. Gy. Makranzy leg. ( HNHM); 3 ♂♂: same data, 30.05.2002 ( HNHM); MEHEDINTI: 1 ♀: Herculane. 22.09- 10.10.1970. T. Palm leg. ( ZML); UKRAINE: VOLYNS’KA OBLAST’: 1 ♀: ‘Polon, Wolhyn Koweł. Th. Krása’ ( NMPC); MOLDOVA: 2 ♂♂: Brostenéi ( IRSNB); GEORGIA: 1 ♀: Adzharo-Imeretsky Mts., 10 km S Sairme. 1100-1850 m a.s.l. 27.06- 02.07.2013. Martynov & Putchkov leg. (cSch); RUSSIA: KARELIA: 1 ♀: Kanadalakshskiy Nature Reserve, west side of Nizhneye Pulongskoe Lake. 03.07.2003. P. Petrov leg. (cR); 4 exs.: ‘Petrosaw[odsk].’, ‘Günther’ ( ZIN); 1 ex.: ‘Jalguba’, ‘Günther’ ( ZIN); 1 ex.: ‘Petrozavodsk Oloneck[aya]. g[uberniya]. G. Yakobson 23.VII.[1]905’ ( ZIN); 2 exs.: ‘Jalguba’ ( ZIN); 1 ♀: ‘Jalguba J. Sahlberg’ ( MZHF); 1 ♀: ‘Saoneskje’, ‘Tolvaja’, ‘B. Poppius’ ( MZHF); 1 ♂: ‘Säämäjärvi Jessoila E.Kangas’ ( MZHF); 2 ♀♀: ‘Perguba. B. Poppius’ ( MZHF); 1 ♀: Kanadalaksha Nature Reserve, west side of Nizhneye Pulongskoye Lake. 03.07.2003. P. Petrov leg. (cR); 1 ♂: Rajajoki. 01.07.1932. R. Elfving leg. ( MZHF); 1 ♂: Sornavala. 19.07.1934. S. Platonoff leg. ( MZHF); 4 ♂♂, 2 ♀♀: same locality. H. Lindberg leg. ( MZHF); 1 ♀: Jaakima. J. Sahlberg leg. ( MZHF); 1 ♂: Paanajärvi. M. Hellén leg. ( MZHF); 2 ♂♂, 4 ♀♀: same locality. 31.07.1936. H. Lindberg leg. ( MZHF); MURMANSK AREA: 2 ♀♀, 1 ex.: ‘Kola, bank of Kola [River] Arkahng[elskaya]. g[uberniya]. Barovskiy [leg.] 22.VIII.[1]926’( ZIN, ZMM); 5 exs.: 12 km SE Poyakonda. 1989. A.O. Benkovskiy leg. ( ZIN); 1 ♀: Poyakonda, biostation of MSU. 12.07.1979. Deshampsia flexuosa . E.M. Veselova leg. (сR); 1 ♂, 2 ♀: Sakharnaya River. 07.1959. S. Tikhomirov leg. ( ZMM); 2 ♀♀: ‘Imandra Envald’ ( MZHF); 1 ♂: ‘Tschavanga. K.M. Levander’ ( MZHF); 1 ♂: ‘Rihpjaur Palmén’ ( MZHF); 2 ♂♂, 2 ♀♀: Yläluostari. S. Platonoff leg. ( MZHF); 1 ♀: Olenitsa. K.M. Levander leg. ( MZHF); 1 ♂: Kuolajärvi, Kutsa. E. Kangas leg. ( MZHF); 1 ♀: ‘Kola Envald’ ( MZHF); 1 ex.: Kola, bank of Kola River. 22.07.1926. V. Barovskiy leg. ( ZIN); ARKHANGELSK AREA: 1 ♂: ‘[Solovetskiye ostrova 3.VII.[18]95. M.N. Rimskiy-Korsakov]’, ‘Solowezki- Inseln Rimsky-Korsakov’ ( MZHF); 1 ♂: ‘Solovetsk’, ‘Envald’ ( MZHF); 1 ex.: ‘Mayak Letniy Orlow [leg.], Onezh[skiy]. u[ezd]. 6 VII.[1]904’ ( ZIN); LENINGRAD AREA: 2 exs.: ‘Yamburg S[anct].-Peterb[urg]. g[uberniya]. V. Barovskiy’ ( ZIN); 1 ♂: Uusikirkko [Polyany]. S. Platonoff leg. ( MZHF); 1 ♂, 1 ♀: Muolaa. E. Palmén leg. ( MZHF); 1 ♀: Konevifs [Konevitsa]. J. Sahlberg leg. ( MZHF); 1 ex.: Sablino. 05.08.1923. L. Bianki leg. ( ZIN); 1 ex.: ‘Fl. Swir’, ‘J. Sahlb[erg].’ ( ZIN); PSKOV AREA: 1 ex.: Remda. 20.06.1901. Sumakov leg. ( ZIN); MOSCOW AREA: 1 ♀: ‘[Moscow] Skhodnya 23.VII.[1]906’ (cR); 2 ♂♂: Morozki st. 01.07.1985. V.B. Semenov leg. (cSh); 1 ex.: Sablino. 05.08.1923. L. Bianki leg. ( ZIN); YAROSLAVL AREA: 1 ex.: ‘N. Kokohyew Iaroslawl’ ( ZIN); STAVROPOL AREA: 1 ♀: Zheleznovodsk. 03.10.1953. I. Telishev leg. ( ZMM); KARACHAY-CHERKESS REPUBLIC: 4 ♂♂: left bank of Daut River, 6 km SW El`brusskiy. 19.06.1993. A. Solodovnikov leg. ( ZMUC); 2 ♂♂, 7 ♀♀: same data, near El`brusskiy. 1400 m a.s.l. ( ZMUC); 1 ♂: slopes of Garallybani Mt., near Dzhenait R. 20.06.1993. 2100-2500 m a.s.l., alpic zone. A. Solodovnikov leg. ( ZMUC); 1 ♂: Medovye Vodopady. 15.06.1994. E.A. Khachikov leg. (cKh); 1 ♂: same sata. 30.07.1994 (cKh); 1 ♂: Teberda ( ZMM); CHECHEN REPUBLIC: 1 ♀: Kobi. 16.07.1911 ( MHNG); KOMI: 1 ex.: ‘Ust`-Ussa Pechorskiy uezd. Arkhangelskaya guberniya. Zhuravskiy. 2 VII [19]05’ ( ZIN); 1 ex.: ‘[6 versts lower Kolva, Ussa [Usa] River, Pechorskiy uezd. Zhuravskiy. 22.VI]’ < Geodromicus plagiatus var. nigrita Mull. B. Poppius det.> ( ZIN); 1 ex.: ‘Ust`-Cylma. Pechorskiy. uezd, Arkhangelskaya guberniya. Zhuravskiy. 18.VI.[19]05’ ( ZIN); 1 ex.: same data. 01.06.1906 ( ZIN); 1 ex.: ‘Bolshezemelskaya tundra, Ibanina River, right tributary of Pechora River. A. Rikhter 12-20.IX.1921 ’ ( ZIN); SVERDLOVSK AREA: 4 ♀♀: Denezhkin Kamen’ Nature Reserve, near Kriv cordon, right side of Kriv Sosvinskiy River near mouth, 60.32710N 59.41849E. 281 m a.s.l., shingle-stony spit along clayish bank. 30.07.2017. O.A. Katana & A.B. Ryvkin leg. (cSh, ZMM); 1 ♂: same data, right side of Sosva River near Kriv cordon, 60.32712N 59.42521E. 286 m a.s.l., stony river bank. 21.07.2017 (cSh); CHELYABINSK AREA: 1 ♂: ‘Urengai: Gory Katav: Ural 1916. Jureček’ ( NMPC); YAMAL-NENEZKIY AUTONOMOUS REGION: 2 exc.: valley of Man`i River, Lesnoy [Forestry] Ural. 22.06.1927. Lyapin & K.F. Flyorov leg. ( ZIN); 1 ♀: same data. 16- 19.06.1927. ( ZIN); 2 ♀♀ Polyarniy Ural, riverbank of Bolshaya Paypuzyna near Polyarniy village. 29.08.1996, D.E. Lomakin leg. ( ZMUC); 1 ♂: ‘[Berezovskiy uezd. Tobolskaya guberniya]’, ‘[Ilovayskiy]’ ( ZMM); KHANTYMANSI AUTONOMOUS REGION: 1 ♀: Khanty-Mansiysk. 05.07.1932. ( ZMM); 2 exs.: environs of Beryozovo. 09.07.1932 ( ZIN); 1 ♀: Sovetskiy District, Malaya Sosva Nature Reserve, right side of Malaya Sosva River near Tunifesh shallow, 61.89675N 64.12277E. 55 m a.s.l., under stones, on pebbles & clay, among grasses on spacious shingle-clayish-sandy spit. 09.07.2017. A.B. Ryvkin leg. ( ZMM); 1 ♀: same District, right side of Malaya Sosva River up-stream of Poryugan River mouth, 61.98127N 63.50690E. 79 m a.s.l., wet small pebbles of shale with quartz intrusions at foot of high steep talus. 11.07.2017. A.B. Ryvkin leg. ( ZMM); 1 ♀: same District, Malaya Sosva Nature Reserve (buffer zone), near Zapadnyi (=Western) cordon, bank of Ay-Yovayugan River, 61.91037N 63.70398E. 72 m a.s.l., small shingles, sand. 02.08.2018. A.B. Ryvkin leg. ( ZMM); KRASNOYARSK TERRITORY: 2 ♂♂, 1 ♀: Turukhanskiy District, Eloguy River, 7-10 km lower mouth of Tynda River. River bank: sand, Carex , grasses, moss, Alnus etc. 19.07.1989. A.B. Ryvkin leg. ( ZMM, cSh); 1 ♀: same District, Yeloguy Refuge. Yeloguy River, 6 km below Tyna River mouth, dried channel of a rill with Alnus sp. and Betula sp. in leaf litter. 27.07.1992. V.B. Semenov leg. ( ZMM); 1 ♂: Turukhanskiy District, Mirnoye, Bolshaya Varlamovka River. 06- 07.08.1979. K. Eskov (cR); 1 ♀: Yermakovskiy District, 4-5 km N Yermakovskoye, Oya River. On silt, in drift, under Salix sp. and Carex spp. 20.08.1984. A.B. Ryvkin leg. (cR); EVENKIA: 1 ♀: Central Siberian Biosphere Reserve, Stolbovaya River: 10 km lower mouth of Dul`kuma River. Under stones near river. 04.07.1993. V.B. Semenov leg. ( ZMM); 1 ♀: Taymura River near mouth of Neptenne River. Forest litter. 01.08.1982. K.Yu. Es`kov leg. (cSh); IRKUTSK AREA: 1 ♂: Katangskiy District, left side of Chona River near mouth of Miringda [=Meringda] River. 521 m a.s.l., thickets of Carex acuta and C. rostrata on wet sand near river. 18- 25.08.2008. A.V. Shavrin & I.V. Enushchenko leg. (cSh); 2 ♂♂: same District, left side of Chona River, 2 km E mouth of Miringda [=Meringda] River, 61º12’417’’N 108º47’238’’E. 280 m a.s.l. 19- 25.08.2008. I.V. Enushchenko leg. (cSh); 1 ♀: Ust`-Kutskiy District, 18 km N Ust`- Kut, valley of Lena River. 391 m a.s.l., under stones near small stream flowing into Lena River. 26- 28.07.2008.A.V. Shavrin & I.V. Enushchenko leg. (cSh); 1 ♂: Nizhneilimskiy District, Nizhne-Ilimsk. 1965. A. Kreslavskiy & V. Zherikhin leg. ( ZMM); 1 ♂: Ziminskiy District, right side of Oka River opposite Bargaday. Under stones, at river. 19- 20.06.1998. A.V. Shavrin leg. (cSh); 1 ♂, 1 ♀: same data, 8 km SE Bargaday, 20.06.1998. A.V. Anishchenko leg. (cSh); 2 ♀♀: Pokoyniki cape, Pokoynickaya River. Under stones near river. 23- 30.06.1995. A.V. Shavrin leg. (cSh); 1 ♀: Slyudyanskiy District, Baikal`sk, valley of Babkha River. 15- 18.07.1992, V. Vladimirov leg. (cS); 1 ♀: ‘Sibir Irkutsk Burguduz, 14.8.1968 T-E. Leiler’ ( SMNH); BURYATIA: 1 ♀: Baikalskiy Nature Reserve, Khamar-Daban Mts., middle flow of Pereemnaya River, winter hut Shum. Valley, mixed forest. 25- 28.07.2009. Yu. Sundukov & L. Sundukova (cSh); 1 ♂, 2 ♀♀: same Mts., Tankhoy, valley of Osinovka Mishikhinskaya River. Under stones near river. 24.07.1995. A.V. Shavrin leg. (cSh); 2 ♂♂, 2 ♀♀: Tunkinskie Mts., Arshan [51°54’ N 102°27’ E]. 23.07- 19.08.1982. V.G. Shilenkov leg. (cSh); YAKUTIA: 1 ♂: Pilka River, right tributary of Lena River, “Pilka”. 06.07.2003. A.I. Averenskiy leg. (cSh); KAZAKHSTAN: EAST KAZAKHSTAN: 21 ♂♂, 24 ♀♀: Altai, Bukhtarma, Archaty. 25.07.1989. V.A. Kastcheev leg. (cSh, ZIN); 2 ♂♂, 7 ♀♀: same data. 29.07.1989 (cSh, ZIN); 3 ♂♂, 3 ♀♀: same data, Dzhambul. 10.08.1988 ( ZIN); 4 ♂♂, 5 ♀♀: same data, Chindagaui. 30.07.1989 (cSh, ZIN); 3 ♂♂, 4 ♀♀: Altai, Rakhmanovskiye Klyuchi, Berel` River. 25.07.1991. V.A. Kastcheev leg. (cSh, ZIN); 1 ♀: Sarymsakty River, Katon-Karagay. 22.07.1989. V.A. Kastcheev leg. ( ZIN); 1 ♂, 1 ♀: environs of Dzhambul, Bukhtarma River. 10.08.1988. V.A. Kastcheev leg. < Geodromicus nigrita Muell. det. Kastcheev> ( ZIN); 2 ♀♀: Tarbagatay, Bazar River. 22.08.1998. V. Kastcheev leg. < Geodromicus nigrita Mull. det. Kastch.> ( ZIN); 2 ♀♀: Zhinishke River, Kyzylaus. 20.06.1988. V. Kastcheev leg. ( ZIN); ALMATY: 1 ♂, 1 ♀: Dzhungar Mts., Tentek River, Kokzhar. 25.08.2006. V.A. Kastcheev leg. (cSh); 1 ♂, 2 ♀♀: same Mts., Oisaz–Chizhe River. 06.07.1990. V.A. Kastcheev leg. ( ZIN); 1 ♂, 4 ♀♀: Ketmen Mts., Shalkydysu River, 17.07.1988. V.A. Kastcheev leg. (cSh); ZHAMBYL: 1 ♀: ‘Aulie Ata SyrDaria’ < Geodromicus a. nigrita Lokay det.> ( SNMB); UZBEKISTAN: 1 ♂: ‘Buchara’, ‘ plagiatus Fbr. det. Luze’ ( HNHM).

Distribution. Geodromicus plagiatus is the most common Palaearctic species of the genus. It is widely distributed in the Palaearctic Region, and known from Europe, the Caucasus, and the mountain regions of Middle Asia eastwards to Lake Baikal ( Fig. 28 ).

Remarks. Staphylinus plagiatus Fabricius, 1798 was originally described based on an unspecified number of syntypes from “Europa borealis”. The studied male ( Figs 1–2 ) from ZMUC is designated here as the lectotype in order to fix the identity of the name. Some authors in the late 19th century considered this species as a variety of G. nigrita (e.g. Reitter 1870).

Lesteva globulicollis Mannerheim, 1830 was originally described from “…lacus Saima in Finlandia orientali…”. The studied holotype ( Figs 3–4 ) from the collection of MZHF is partly damaged. It seems to have been removed from the original pin and dissected ( Fig. 3 ). For a long time, this taxon was considered as a distinct species (e.g. Thomson 1861, Redtenbacher 1874, Ganglbauer 1895). It was one of the examples of species with confusing interpretations, despite the fact that it was synonymized with G. plagiatus by Erichson (1840). Geodromicus globulicollis was considered as a variety of G. brevicollis Fauvel, 1875 by Neumann (1907), G. curtipennis Fauvel, 1871 by Bernhauer & Schubert (1910) and Scheerpeltz (1933), and G. lituratus Kraatz, 1857 (now a synonym of G. kunzei ( Heer, 1839)) by Luze (1903). Khnzorian (1962) cited Caucasian G. constricticollis Eppelsheim, 1890 as a subspecies of G. globulicollis . Several species were cited as synonyms of G. globulicollis : G. brevicollis by Bernhauer & Schubert (1910), G. curtipennis by Ganglbauer (1895), G. kunzei ( Heer 1839) by Kraatz (1857) and Ganglbauer (1895), G. lituratus by Bernhauer & Schubert (1910), G. longipes ( Mannerheim, 1830) by Kraatz (1857), Ganglbauer (1895) and others, and others, G. puncticollis (Weise, 1875) by Eppelsheim (1887), Trichodromeus flavipes ( Motschulsky, 1860) by Gemminger & Harold (1868) and T. transversus ( Motschulsky, 1860) by Fauvel (1871). As a result of such confusing interpretations, many faunistic records require clarification, e.g., the record of G. plagiatus a. nigrita from Tiflis (=Tbilisi) by Eichler (1930).

Anthophagus verticalis Say, 1831 was originally described based on an unspecified number of syntypes from “ Pennsylvania ” ( USA) and compared with Anthophagus brunneus Say, 1823 . Later it was recorded from “Lake Superior” by LeConte (1850). Without studying type material Fauvel (1878) synonymized it with the European G. nigrita . Ganglbauer (1895) and subsequent compilers cited A. verticalis as a synonym of G. plagiatus . This species, in turn, was synonymized by Casey (1894) with the North American G. brunneus ( Say, 1823) . This synonymization was overlooked by compilers who synonymized this species with other European taxa. I included this name to the list of synonyms of G. plagiatus with the mark <?!>, because it seems that “ A. verticalis ” is not conspecific with the Palaearctic G. plagiatus and the type material should be studied. In general, all Nearctic species of Geodromicus require revision. Based on my data, Geodromicus brunneus , with which Casey synonymized A. verticalis , includes two different and not closely related species from Canada and the USA.

Lesteva canaliculata Stephens, 1831 was originally described from Britain based on an unspecified number of syntypes: “Taken in Scotland by Mr. Hooker, and also found in Cumberland: not common. Near Edinburgh ”. Waterhouse (1858) synonymized it with G. globulicollis . Subsequent authors considered L. canaliculata as a synonym of G. plagiatus (see Herman 2001). According to Pope (1977), G. plagiatus does not occur in Great Britain (see also Duff 2012) and old records of this species are doubtful. The type locality of L. canaliculata was not included in the map of G. plagiatus ( Fig. 28 ).

Geodromicus anthracinus Brisout de Barneville, 1863 was originally described based on an unspecified number of syntypes from “Mont-Louis” (Southeast France) and was compared with G. plagiatus . Fauvel (1871 [= 1872]), Ganglbauer (1895) and subsequent authors considered G. anthracinus as a synonym of G. plagiatus .

Geodromicus subnigritus was originally described as an aberration of G. plagiatus without citing of the type locality. It was recorded by Palm (1961) from Sweden. Zanetti (1987) synonymized it with G. plagiatus . I studied two specimens with additional labels “ Typus ” and “paratypoid” from MHNG. Both specimens are from Austria. In any case, this name is unavailable (see details in Herman 2001).

Geodromicus pyrenaeus Jarrige, 1963 was originally described from the Pyrenees, France (“Gave d’Holçarté, Larrau, 600 m, Basses Pyrénées”).A study of the type material did not reveal any external and internal morphological differences between this material and other specimens of G. plagiatus . Consequently , G. pyrenaeus is synonymized with it.

Geodromicus lapponicus A. Bordoni, 1984 was originally described based on an immature male with a strongly deformed median lobe ( Bordoni 1984: fig. III.1) without exact locality from “Lapponia”. A study of the holotype revealed that it belongs to G. plagiatus . Schülke & Smetana (2015) cited this name as a nomen dubium.

Geodromicus klimai A. Bordoni, 1984 was originally described based on five specimens without exact locality from “Caucaso”. I studied the holotype and did not find any morphological differences suggesting that it should represent a distinct species. The illustrations of the apical portion of the median lobe provided by Bordoni (1984: fig. III.3) is somewhat similar to that of G. nigrita ( Figs. 36–40 ), but the aedeagus of the holotype is slightly narrower. Thus, G. klimai is synonymized with G. plagiatus .

Geodromicus schoenmanni Bordoni, 1984 was originally described based on six specimens without exact locality from “Caucaso”. I studied the holotype ( Fig. 6 ) and three paratypes and found no clear morphological differences between them and other specimens of G. plagiatus . The shape of the aedeagus ( Fig. 18 ) is similar to specimens from other localities. In consequence, it is synonymized with G. plagiatus .

Regarding the habitus, punctation of the forebody, shape of the elytra (slightly broadened apicad), and general shape of the aedeagus, G. plagiatus is very similar to G. nigrita (see below). According to European keys (e.g. Zanetti 2011), G. plagiatus can be distinguished from G. nigrita by paler legs, sometimes by the presence of a broad yellow to reddish spot on each elytron, and the shape of the apical portion of the aedeagus. The coloration of G. plagiatus varies from yellow-brown to dark-brown, and legs are yellow to dark-brown ( Fig. 5 ). The presence of a broad spot on each elytron occurs in several species of the genus. This feature does not allow a reliable distinction of G. plagiatus and G. nigrita . I observed the presence of similar spots in several Palaearctic species, which usually have unicolorous elytra: G. brevicollis , G. bodemeyeri Bernhauer, 1902 , G. convexicollis Luze, 1903 , G. latiusculus Eppelsheim, 1878 , etc. Many authors of the last decades have interpreted the presence of elytral spots as a significant feature in their diagnoses, which has resulted in various interpretations of different taxa and wrong synonymizations. Considerable interspecific variability of G. plagiatus was observed in body size (from 3.9 to 6.7 mm), punctation of the head and pronotum (finer and sparser to coarser and denser), the proportions of the pronotum, the shape of the apical portion of the median lobe, and the length of the parameres ( Figs. 12–27 ). The shape of the apical portion of the median lobe is one of the main features for the distinction from related species. The apex is narrower in G. plagiatus and distinctly broader in G. nigrita ( Figs. 36–40 ). The aedeagus of G. plagiatus were figured by Palm (1948: fig. 183), Lohse (1958: Fig. 2 ), Fagel (1958: Fig. 1 ), Smetana (1959: Fig. 8 ), Lohse (1964: Fig. 2 ), Tóth (1982: Fig. F), Bordoni (1984: Fig. 1–2 ), and Zanetti (1987: Figs 96e, 96f). The figure of the aedeagus provided by Smetana (1959: fig. 11) as G. globulicollis refers to G. kunzei . I decided to give figures of the aedeagi of specimens from different localities to show variability of their shapes ( Fig. 12–27 ). In general, the shape of the aedeagus is slightly variable: the preapical portion of the median lobe can be somewhat broader ( Fig. 19 ) or narrower ( Fig. 15 ), with subacute ( Figs. 12–15, 17 , 21 ) or acute ( Figs. 16, 18–20, 22–26 ) apex, and the parameres may not extend to, or slightly extend beyond, the apex of the median lobe.

Herman (2001) considered G. plagiatus as a species with a Holarctic distribution. It was recorded for “Peninsulo Ibérica” ( Gamarra & Outerelo 2008). Ryabukhin (1999) recorded it from the Magadan area (“Upper Kolyma: environs of Orotukan and 30 km NE of Spornoye”), but this record requires clarification. The species is here recorded from Cataluña province of Spain, Moldova, Georgia, Uzbekistan, and several regions of Russia (Pskov Area, Stavropol Area, Karachay-Cherkess Republic, Chechen Republic, Sverdlovsk Area, Khanty-Mansi Autonomous Region, Krasnoyarsk Territory, Evenkia and Yakutia) for the first time.

Figures

FIGURE 28. Distribution of Geodromicus plagiatus in the Palaearctic Region. Blue circles: specimens studied by author; green circles: previously published records, specimens not seen by author.

FIGURES 1–11. Habitus (Figs. 1, 3, 5–11) and type labels (Figs. 2, 4) of species of Geodromicus: 1–2–Staphylinus plagiatus (lectotype), 3–4–Lesteva globulicollis (holotype), 5–G. plagiatus (Russia: Buryatia, Arshan), 6–G. schoenmanni (holotype), 7–G. pyrenaeus (paratype), 8–G. corsicus (Corsica, Vizzavona), 9–G. nigrita (Czech Republic: Čertovo), 10–G. pedemontanus (France, Boreon), 11–G. suturalis (Switzerland: Rhone). Scale bars: 1.0 mm.

FIGURES 12–19. Aedeagus (parameral view) of Geodromicus plagiatus: 12–France, Bious Artiguer, 13–France, Gavarnie, 14–Czech Republic, Nový Svĕt, 15–Sweden, Abisko, 16–Sweden, Vistbäcken, 17–Russia, Karachay-Cherkessia, Elbrus, 18– ‘Kaukasus’ (paratype of G. schoenmanni), 19–Russia, Murmansk Area, Yläluostari. Scale bars: 0.1 mm.

FIGURES 20–27. Aedeagus of Geodromicus plagiatus from Russia (Figs. 20–24, 26–27) and Kazakhstan (Fig. 25) (Figs. 20–26: parameral view, Fig. 27: lateral view): 20–Moscow Area, Morozki, 21–Sverdlovsk Area, Sosva River, 22–Krasnoyarsk Territory, Yaloguy River, 23–Irkutsk Area, Bargaday, 24–Buryatia, Arshan, 25–Almaty, Ketmen Mts., 26–27–Irkutsk Area, Miringda River. Scale bars: 0.1 mm.

FIGURES 36–40.Aedeagus of Geodromicus nigrita (Figs. 36–39: parameral view, Fig. 40: lateral view): 36–Spain, La Granja, 37–Spain, Sierra Nevada, 38–Austria, Stoder, 39–40–UK, Ribbiesdale. Scale bars: 0.1 mm.