Lamproclasiopa lapaz, Costa, Daniel N. R., Mathis, Wayne N. & Marinoni, Luciane, 2016

Costa, Daniel N. R., Mathis, Wayne N. & Marinoni, Luciane, 2016, A revision of the shore-fly genus Lamproclasiopa Hendel (Diptera, Ephydridae), ZooKeys 631, pp. 1-99 : 29-33

Taxon classification Animalia Diptera Ephydridae

Lamproclasiopa lapaz sp. n. Figs 31-35, 36

Diagnosis.

This species is distinguished from congeners by the following combination of characters: Moderately small shore flies, body length 2.40-2.97 mm; generally a shiny black species. Head: Frontal and facial microtomentum sexually dimorphic; male with dense and extensive microtomentum on the frons, also within ocellar triangle, anterior laterals of frons bare, shiny black, Female frons with broad, transverse stripe of microtomentum on the center; male mesonotum with microtomentum on anterior third. Antenna black, with dense microtomentum especially evident on basal flagellomere laterally; arista bearing 3-4 dorsal rays (usually 4). Face with moderately deep antennal grooves on dorsal half, shallowly angulate in lateral view, vortex of angle at midheight near dorsal facial seta, ventral half of face receded, male facial microtomentum generally dense, golden brown dorsally, becoming more silvery ventrally, female face most bare, with silvery microtometum at the ventral portion of face and at base of facial setae and adjacent to parafacial; parafacial and anterior half of gena densely microtomentose in male, in female with thin area microtomentose at anterior and ventral margins of eye, otherwise bare, shiny; gena very high, gena-to-eye ratio 0.42-0.54; posterior margin of gena at merger with lateral margin of postgenal sharply angulate. Thorax: Mesonotum shiny black, male mesonotum with microtomentum on anterior third, with a thin lateral extension at level of suture and extended along posterior margin of notopleuron; lateral to microtomentose band mostly bare, shiny except for microtomentose anterior surface of postpronotum and ventral margin of notopleuron; presutural supra-alar seta well developed; pleural region generally bare, shiny black. Wing hyaline to faintly infumate, faintly tannish, lacking any pattern or markings. Costal vein ratio 0.40-0.51; M vein ratio 0.69-0.83. Coxae black, shiny; forecoxa with some lateral areas microtomentose but lacking vertical microstriae; femora and tibiae black; forefemur with 4-5 stout, peg-like setae on apical third along posteroventral margin; basal 2-3 tarsomeres yellow, apical 2-3 brownish black to dark brown. Halter with base black, knob whitish yellow. Abdomen: Generally shiny black; male tergite 5 truncate apically. Male Terminalia (Figs 31-35): Epandrium in posterior view (Fig. 31) with dorsal half transversely rectangular, lateral margin shallowly convex, dorsal margin broadly truncate, very thin above cercal cavity, ventral half thinner than dorsal half, demarcation sharply angulate, thereafter ventral extensions almost parallel sided, ventral margin broadly bilobed with moderately deep, thin, incision, setulae clumped, at ventral margin, at beginning of ventral half and at 2 sites along dorsal margin, in lateral view (Fig. 32) with posterior portion linear, thinnest dorsally and subventrally, thereafter ventrally enlarged, clavate, with well-developed anterior, hook-like extension, hook angulate rather than rounded, bearing setulae at vortices of angles; cerci in posterior view (Fig. 32) narrow, elongate, rod-like, slightly wider dorsally, apparently fused ventrally with ventral margin of cercal cavity, in lateral view (Fig. 33) narrowly lunate; aedeagus in lateral view (Fig. 35) as 2 structures of differing lengths, elongate basiphallus curved, irregularly tapered, base of basiphallus T-shaped, bar formed by pointed, lateral projections, apex of basiphallus acutely pointed and more curved, distiphallus less than half length of basiphallus, shallowly curved, otherwise rod-like, in ventral view (Fig. 34) with basiphallus as a thick, inverted Y, incised gap narrowly and deeply U-shaped with a heart-shaped extension at base, distiphallus with base within apical gap of basiphallus, narrow, straight, rod-like; phallapodeme in lateral view (Fig. 35) C-shaped, each arm expanded apically, in ventral view as 2 stacked, moderately broad, short arrowheads; gonite in lateral view irregularly clavate, narrow, elongate, straight, rod-like, in ventral view (Fig. 34) shorter than gonite in lateral view, rod-like; hypandrium in lateral view (Fig. 35) thin, elongate, irregularly clavate basally with midlength, short projections, in ventral view (Fig. 34) as 2 irregular, almost parallel, rectangular sclerites, wider anteriorly than posteriorly, with a W-shaped base with narrow arms extended posteriorly and slightly laterally, and 2 short medial bumps along base.

Type material.

The holotype male of Lamproclasiopa lapaz is labeled "BOLIVIA. La Paz: La Paz (6 km NE; 16°25.7'S, 68°04.3'W; 4130m), 19 Mar 2001[,] Wayne N. Mathis/ USNM ENT 00119994 [plastic bar code label]/HOLOTYPE ♂ Lamproclasiopa lapaz Costa, Mathis & Marinoni USNM [red]." The holotype is double mounted (minuten pin in a block of plastic), is in good condition (abdomen removed, dissected, parts in an attached microvial), and is deposited in the USNM. Two female paratypes bear the following label data: Bolivia. La Paz (NE; 16°27.4'S, 68°06'W; 3940m), 19 Mar 2001, W. N. Mathis (2♀; USNM).

Type locality.

BOLIVIA. La Paz: La Paz (6 km NE; 16°25.7'S, 68°04.3'W; 4130m).

Distribution

(Fig. 36). Neotropical: Bolivia (La Paz).

Etymology.

The species epithet, lapaz , refers to the capital of Bolivia, La Paz, where the type series was collected. La Paz is Spanish for peace, which we embrace and recommend to all.

Remarks.

This species is very similar both of the other species of the polita group, especially Lamproclasiopa auritunica , but is distinguished from these two species as follows: Female frons with broad, transverse stripe on anterior half (female frons in Lamproclasiopa auritunica is mostly bare, shiny black); male mesonotum with microtomentum on anterior third only (male mesonotum in Lamproclasiopa auritunica has a broad longitudinal band over entire mesonotal length, although it is weaker posteriorly). Shapes of structures of the male terminalia are also diagnostic.