Strobila leiodon Hutton, 1882

Brook, Fred J., Kennedy, Martyn, King, Tania M., Ridden, Johnathon, Shaw, Matthew D. & Spencer, Hamish G., 2020, Catalogue of New Zealand land, freshwater and estuarine molluscan taxa named by Frederick Wollaston Hutton between 1879 and 1904, Zootaxa 4865 (1), pp. 1-73 : 29

Strobila leiodon Hutton, 1882

Pl. 3, fig. D

Hutton, 1882. New Zealand Journal of Science, 1: 281.

Type material. Lectotype (designated here), CMNZ 59.3, and paralectotypes, CMNZ M59 (6), CMNZ M9923 [ex M59] (2) (dry shells). Details in the CMNZ molluscan catalogue indicate that lots M1065 [ZS 723] and M12776 [ex M1065] are also primary type material of Strobila leiodon Hutton, 1882 (see below), with the former lot being listed by Freeman et al. (1997: 31). However, neither of these lots were found during a search of the CMNZ molluscan collection in 2017. The collection at CMNZ has a radula mounted on a glass slide with the label details ‘ Huttonella leioda, Greymouth, XVI p. 166’, in Hutton’s handwriting (i.e., CMNZ 2017.17.105), which is probably primary type material (see descriptions of radula by Hutton 1883d: 136, 1884b: 166).

Label details. CMNZ M59—‘69. Strobila leiodon Hutton , Greymouth’, pillbox label in Hutton’s handwriting.

CMNZ molluscan catalogue details. M1065—‘ Endodonta leiodon Hutton, Greymouth (old number ZS 723)’.

Type locality. ‘Greymouth (R. Helms)’ ( Hutton 1882p: 281, 1883d: 136).

Previous illustrations of type material. Radula teeth illustrated by Hutton (1884b: pl. 9, fig. P) probably from type material; Pilsbry (1892 [in 1892–1893]: 87, pl. 24, figs. 32–35—‘drawn by Mr. Suter from the type specimens’); Suter (1915: pl. 9, figs. 16, a–c—possibly type material).

Remarks. Hutton submitted a description of Strobila leiodon to the Transactions and Proceedings of the New Zealand Institute issue for 1882, but publication was delayed until May 1883 ( Hutton 1883d: 135), and was preempted by a brief description in an account of a meeting of the Philosophical Institute of Canterbury ( Hutton 1882p: 281). A description by Hutton (1884b: 166, pl. 9, fig. P) of a radula of leiodon was probably based on the putative type material in CMNZ 2017.17.105 mentioned above. Hutton’s descriptions of S. leiodon were based on more than one specimen, and he did not designate a holotype. Climo (1969: 194) incorrectly stated that the ‘holotype’ was in Canterbury Museum, but this is not a valid lectotype fixation according to either Art. 74.5 or Art. 74.6 of the ICZN (1999) Code, because he was aware that the type material consisted of more than one specimen, and did not explicitly indicate that he was selecting a particular specimen to serve as the name-bearing type. Strobila leiodon Hutton, 1882 is the type species of Ptychodon Ancey, 1888 , by original designation. It is also the type species of Huttonella Suter, 1890 (non Huttonella Pfeiffer, 1856 ), by subsequent designation of Schileyko (2001: 937), and hence also of the replacement genus name Maoriana Suter, 1891 (see ICZN Article 67.8). The two last-mentioned genus names were treated as junior synonyms of Ptychodon by Schileyko (2001: 937).

Current taxonomy. Ptychodon leiodon (Hutton, 1882) — Ancey (1888: 372), Iredale (1915: 481), Climo (1978a: 181, 1989: 623), Spencer et al. (2009: 216).

Distribution. New Zealand; western South Island, predominantly in the northern part between Golden Bay and Hokitika, with scattered records south to Hollyford Valley, Fiordland ( Climo 1989: 625, fig. 22; AIM and NMNZ collection records).