Succinoraphidia radioni, 2019

Perkovsky, Evgeny E. & Makarkin, Vladimir N., 2019, A new species of Succinoraphidia Aspöck & Aspöck, 2004 (Raphidioptera: Raphidiidae) from the late Eocene Rovno amber, with venation characteristics of the genus, Zootaxa 4576 (3), pp. 570-580 : 572-574

Succinoraphidia radioni sp. nov.

Figs 1–4

Diagnosis. In both fore- and hind wings, ScP relatively long, length between termination of ScP and proximal end of pterostigma twice shorter than length of pterostigma [this length slightly longer than length of pterostigma in S. baltica ]; pterostigma distinctively paler in proximal part, darker in distal part [somewhat darker proximally in S. exhibens ]; in hind wing, anterior trace of MP with three pectinate branches [two branches in S. exhibens ].

Description. Female. Body poorly discernible. Prothorax relatively broad. Ovipositor long, with oval gonostyli ( Fig. 3 ).

Forewing oval, 8.7 mm long, 2.7 mm wide. Costal space moderately dilated at 1/5 wing length; all four subcostal veinlets simple, widely spaced (additional incomplete veinlet between two basal veinlets in right wing). ScP relatively long, terminating on costal margin at about 0.55 wing length; length between termination of ScP and proximal end of pterostigma twice shorter than length of pterostigma. Subcostal space broad, with two crossveins: basal crossvein 1scp-r located distad seeming origin of M; distal crossvein 2scp-r at proximal end of pterostigma. Pterostigma elongate, paler proximad proximal branch of RP (light-brown), darker distad (brown). Anterior trace of RA not forked distally, strongly bent posteriorly after origin of distal branch, terminating at margin before wing apex; with two branch (veinlet): proximal branch located within pterostigma (part of pterostigma proximad this branch longer than distad); distal one nearly straight, located at distal end of pterostigma. RA space (between RA, RP) with two crossveins: 2ra-rp long, located slightly proximad proximal end of pterostigma; 3ra-rp shorter, located well distad end of pterostigma. RP originates far from wing base (at about 0.48 complete wing length), with two branches: RP1 deeply forked at outer gradate series of crossveins; RP2 simple. Anterior trace of RP not forked distally. One crossvein between branches of RP as part of outer gradate series (irp), connecting anterior branch of RP1, RP2 at its origin. Three crossveins between RP, MA: 1r-m, 2r-m connect anterior trace of RP, MA; 3r-m connects RP1 and anterior branch of MA. M appear fused basally with R for considerable distance, then actually fused with CuA for short distance; forked distad origin of RP. MA once deeply forked. MP fused with CuA for considerable distance, then pectinately branched, with two simple branches. Two intramedian crossveins form two doi. Cu dividing into CuA, CuP rather far from wing base. CuA crossvein-like before fusion with M, perpendicular to Cu, then fused with M for very short distance, not forked distally. CuP simple, aligned with Cu and nearly parallel to distal part of CuA. Crossvein between CuA, CuP (icu) long; crossvein between CuP, AA1 short. Anal veins not forked distally; no crossvein between these. AA2, AA3 fused basally; then AA2 bent to AA1 and fused with it for short distance; free distal part of AA3 short.

Hind wing ca. 7.5 mm long, 2.6 mm wide. Costal space narrow; subcostal veinlets poorly discernible. ScP relatively long, terminating on costal margin at about 0.63 wing length; length between termination of ScP and proximal end of pterostigma nearly twice shorter than length of pterostigma. Subcostal space with one distal crossvein 2scp-r at proximal end of pterostigma. Pterostigma elongate, slightly lighter than in forewing; paler proximad proximal branch of RP, darker distad. Anterior trace of RA not forked distally, strongly bent posteriorly after origin of distal branch, terminating at margin before wing apex (slightly distad than in forewing); with two branch (veinlet): proximal branch located within pterostigma (part of pterostigma proximad this branch longer than distad); distal one nearly straight, located at distal end of pterostigma. RA space with three crossveins: 1ra-rp located nearly opposite to 2r-m; 2ra-rp located somewhat proximad proximal end of pterostigma; short 3ra-rp located well distad termination of pterostigma, connecting anterior traces of RA and RP. RP originates relatively close to wing base (at ca. 0.35 complete wing length), with two branches. RP1 deeply forked outer gradate series of crossveins; RP2 simple. One crossvein between branches of RP as part of outer gradate series. Three crossveins between RP, M/MA; 1r-m long, slightly sinuate, connecting anterior trace of RP and M near its seemingly origin; 2r-m connects RP1 and anterior trace of MA; 3r-m as part of outer gradate series connecting RP1 before fork and anterior branch of MA. M appears fused basally with R, forked into MA, MP distad well origin of RP. MA deeply forked distally; MP pectinately branched, with three branches. Two crossveins between MA and MP (1im, 2im) form two cells (1doi, 2doi), nearly equal in length; length/width ratio of 2doi 1.7 (left wing) to 2.1 (right wing). Two crossveins between M and Cu: basal 1m-cu moderately long, only slightly oblique, located near wing base; 2m-cu long, connecting MP and anterior branch of CuA. CuA deeply forked. CuP simple. One long crossvein between CuA, CuP (icu), located far distad 1m-cu. Cu and AA1 fully separate; one short crossvein connecting these located between 1m-cu, 2m-cu. AA1, AA2 fused proximally; their distal (free) parts simple. AA3 short, simple (poorly discernible). One long crossvein between AA1+AA2 and AA3.

Etymology. From the first name of Radion F. Khomich, father of the collector of the specimen.

Type material. Holotype: Specimen No. SIZK L-105, collected by Nikolai R. Khomich and currently deposited in his collection, but will be ultimately deposited in SIZK. A poorly-preserved, slightly incomplete female.

Locality and horizon. Rovno amber ( Voronki in Vladimirets District of Rovno Region, Ukraine); late Eocene .

Remarks. The new species easily differs from S. baltica by the longer ScP. Unfortunately, the venation of S. exhibens is incompletely described. However, the photograph of the holotype and the drawing of the basal part of the hind wing (see Aspöck & Aspöck 2004: Figs 1 , 5 ) show that the venation of S. exhibens differs from that of the new species by the number of MP branches and the presence of additional long crossvein between MP and CuA in the hind wing, as well as by the coloration of the pterostigma (this is “mainly yellowish, proximally somewhat darker” in S. exhibens: Aspöck & Aspöck 2004, p. 12 ).

Figures

FIGURE 1. Succinoraphidia radioni sp. nov., holotype SIZK L-105 as preserved. A, left view; B, dorsal view. Scale bars = 2 mm.

FIGURE 2. Succinoraphidia radioni sp. nov., holotype SIZK L-105, right wings. Scale bar = 2 mm.

FIGURE 3. Succinoraphidia radioni sp. nov., holotype SIZK L-105, apex of ovipositor. A, ventral view; B, latero-caudal view. gs9, gonistyli of 9th segment; gx9, gonocoxites of 9th segment. Scale bars = 0.1 mm.

FIGURE 4. Wing venation of Succinoraphidia radioni sp. nov., holotype SIZK L-105. A, left forewing; B, right forewing; C, left hind wing; D, right hind wing (A and C converted to standard view, with apex to the right). Fused veins are shown by red lines. Scale bar = 2 mm (all to scale).

FIGURE 5. Succinoraphidia baltica (Carpenter, 1957), holotype MCZ 5122. A, ventral view; B, dorsal view. Scale bars = 2 mm.