Jakliella papuana, Horák, 2010

Horák, Jan, 2010, Jakliella and Mehlia, new genera from Indonesia and Malaysia, with description of four new species (Coleoptera: Mordellidae), Acta Entomologica Musei Nationalis Pragae 50 (2), pp. 547-561 : 558-560

Jakliella papuana sp. nov.

( Figs. 36–46 )

Type material. HOLOTYPE: J, ‘ Indonesia, West Papua [= Irian Jaya], Arfak Mts, Monokvari distr., cca 20 km S of Warmere, Dubbei env., 21.i.–8.ii.2008, St. Jakl leg.’ ( NMPC) . ALLOTYPE: ♀, the same data ( JHPC) .

Description. Male. Body black, 3 mm long from tips of mandibles to apex of elytra and 3.6 mm long from tips of mandibles to apex of hypopygium. Anteclypeus, anterior border of labrum, galea, maxillary palpi and four basal antennomeres yellow brown to red brown. Legs black, only anterior femora indistinctly black brown. Terminal spurs of metatibia black. Pubescence of dorsal surface black with white sutural stripe on elytra ( Fig. 36 ). Underside with black pubescence, basal portions of abdominal ventrites, hypopygium and pygidium covered with silvery hairs.

Head flatly convex, shining and sparsely punctate, wider than long (13: 11), only moderately prolonged anteriorly. Galea short, narrow, with yellow cirrose outgrowths at distal end. Eyes ( Fig. 37 ) of usual size, almost rhomboid-shaped with rounded angles, finely facetted and glabrous. Neither temples nor temporal angles developed. Maxillary palpi ( Fig. 38 ) robust, ratio of its length to length of antenna equal to 7: 12; palpomere 2 only indistinctly wider than palpomere 3; terminal palpomere broadly securiform, with inner angle situated at midlength. Antennae short ( Fig. 40 ), nearly as long as width of head; antennomere 3 approximately 0.7 times as wide and as long as antennomere 2 and 0.75 times as long and indistinctly narrower than antennomere 4; antennomere 5 about 1.2 times as long and 1.25 times as wide as antennomere 4; antennomeres 5–10 as long as wide; terminal antennomere oblong oval, about 1.3 times as long as penultimate one.

Pronotum flatly vaulted, widest in basal third, distinctly wider than long (15: 11). Anterior angles hardly visible from above, anterior margin almost semicircular with indistinct neck-shaped protuberance. Sides straight in lateral view, posterior angles obtuse with widely rounded tips. Punctation dense and rasp-like.

Scutellum broadly quadrangular with black pubescence.

Elytra 2.25 times as long as their combined width at humeri, gradually and regularly narrowed posteriorly, their tips separately rounded. Punctation very coarse, rasp-like, especialy around scutellum.

Pygidium short, 1.4 times as long as hypopygium and reaching one fourth of length of elytra, rather broadly conical, with broad ring of white hairs at base.

Metepisterna long and narrow, distinctly wider than elytral epipleura in humeral portion, rounded at their posterior end.

Anterior tibiae ( Fig. 42 ) distinctly curved inwards, at base not dilated and without long hairs. Anterior tarsi distinctly narrower than protibiae, two basal anterior tarsomeres subequal; tarsomere 3 approximately 0.75 times as long as and indistinctly narrower than tarsomere 2; tarsomere 4 about 0.75 times as long as and indistinctly narrower than tarsomere 3, truncate at apex; terminal tarsomere cylindrical, 2.0 times as long as preceding one. Middle tibiae 0.85 times as long as middle tarsi. Posterior tibia with only one short apical ridge reaching one fourth of width of tibia; posterior tarsomeres without ridges. Outer terminal spur of metatibia reaching one third of the inner one.

Male genitalia as figured ( Figs. 43–45 ), shape of internal urosternite 8 as in Fig. 46 .

Sexual dimorphism. Female (allotype). Body only indistinctly more robust than in male, 3.4 mm long from tips of mandibles to apex of pygidium. Antennae extremely short ( Fig. 41 ); antennomeres 5–10 distinctly wider than long; terminal antennomere quadrate. Maxillary palpomeres 2 and 3 black brown; terminal palpomere black, unusually large and swollen ( Fig. 39 ). Maxillary palpus about 0.65 times as long as antenna. Pubescence of dorsal surface black with three indistinct whitish bands on pronotal disc and sutural stripe running from apex of scutellum to apex of elytra.

Etymology. The specific name refers to the distribution of the new species in West Papua (part of the former Irian Jaya).

Distribution. Indonesia: West Papua.

Figures

Figs. 36–46. Jakliella papuana sp. nov. 36 – general view; 37 – eye; 38 – maxillary palpus; 39 – maxillary palpus; 40 – antenna; 41 – antenna; 42 – anterior tibia and tarsus; 43 – paramere; 44 – phallobasis; 45 – apical part of penis; 46 – 8 internal sternite. 36–38, 40, 42–46 – holotype (male); 39, 41 – allotype (female). Scale: a – 38–41; b – 46; c – 37; d – 42, 44–45; e – 43; f – 36.