Nipponaphaenops (Tanakaphaenops), Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün, 2022

Karaoğlan, Beliz Bahar, Yekedüz, Emre, Yazgan, Satı Coşkun, Mocan, Eda Eylemer, Köksoy, Elif Berna, Yaşar, Hatime Arzu, Şenler, Filiz Çay, Utkan, Güngör, Demirkazık, Ahmet, Akbulut, Hakan & Ürün, Yüksel, 2022, Contribution to the Knowledge of the Rakantrechus Complex (Coleoptera, Carabidae, Trechini), with Description of a New Subgenus and a New Species of the Genus Nipponaphaenops S. Uéno, 1971 from Northwestern Shikoku, Western Japan, Bulletin of the National Museum of Nature and Science. Series A, Zoology (Zoology) 48 (3), pp. 119-138 : 131-132

publication ID

https://doi.org/ 10.50826/bnmnszool.48.3_119

persistent identifier

https://treatment.plazi.org/id/A95B8783-390A-FFE7-FD47-FD72FB0EC3BF

treatment provided by

Felipe

scientific name

Nipponaphaenops (Tanakaphaenops)
status

subgen. nov.

Subgenus Tanakaphaenops subgen. nov.

urn:lsid:zoobank.org:act:9F3207A6-8680-471F-A467-5279EDBBE3DC

Type-species: Nipponaphaenops kojii sp. nov.

Diagnostic characters. Discriminated from the nominate subgenus by the following character states; body size smaller (BL: 5.39 mm); head smaller, shorter (PW/HW 1.30, PL/HL 1.50, HW/ HL 0.98); frontal furrows complete; tempora to genae with a temporal seta in addition to a genal seta; penultimate labial palpomere about 1.05 times longer than ultimate one, with two subapical setae on the external face of the former usual in size ( Fig. 1n View Fig ); antennae shorter, stouter, less than a half longer than elytra; pronotum with hind angles obtuse at the tip; posterior latero-marginal setae of pronotum present; elytra shorter (EL/EW 1.39), with prehumeral borders shorter and more straight; pore 1 of the marginal umbilicate series of elytra more distant from marginal gutter and closer to elytral base (U1 15.8), pore 4 of the series closer to elytral base (U4 29.2); elytral discal seta of the external series more externally (in stria 5), more posteriorly (basal 11/17) positioned; legs shorter and stouter; ventrites 4–6 each with a pair of paramedian setae ( Fig. 2k View Fig )

Description. Body size small for the genus.

Chaetotaxy of head as in the nominate subgenus except that genae with a temporal seta; both two pairs of latero-marginal setae of pronotum present; elytral discal and preapical setae as in the nominate subgenus; pore 1 of the humeral set of marginal umbilicate series of elytra postero-dorsally translocated, positioned on stria 7 and close to the level of the pore 2 of the series, and pore 4 of the series widely distant from pore 3 ( Fig. 3d View Fig ). Ventrite 7 of female with two pairs of apical setae ( Fig. 2k View Fig ).

Head elongate subquadrate; frontal furrows complete, deep in most part, though becoming superficial behind the level of posterior supraorbital pores; labial suture nearly completely vanished; mentum tooth relatively long and slender, sharply bifid at apex ( Fig. 2e View Fig ); right mandible ( Fig. 1i View Fig ) with a tricuspid retinacular tooth (which is supposedly formed by the union of premolar and primarily bicuspid retinaculum) whose median cusp is rather reduced (cf. Deuve et al., 2016, p. 380).

Pronotum elongate subcordate, wider than head, obviously longer than wide, with front angles rounded but advanced, with hind angles as a whole subrectangular though obtuse at the tip.

Elytra large, oval, with basal parts strongly rectilinearly constricted basad, with prehumeral marginations becoming evanescent anteriad a short way before shoulders; baso-lateral areas of elytral disc just before shoulders nearly vertically deflexed ventrad, forming oblique longitudinal carinae consisted of basal parts of intervals 6–7, with the outlines of prehumeral portion of elytra barely visible from above beneath these carinae ( Fig. 3d View Fig ); basal portions of intervals 2–5 more or less depressed, while basal parts of interval 1 including scutellar area left undepressed as in the nominate subgenus; inclined face of basal peduncle rather narrowly flat, embraced in gentle slope, which is bordered posteriorly by the obtuse transverse carinae formed by the basalmost portion of elytral disc ( Figs. 3c–d View Fig ).

Legs long, but relatively stout.

Male characters. Unknown.

Taxonomic notes. This subgenus is presently monotypic. In this subgenus (as far as known from the type species), the position of the first pore of the marginal umbilicate series of elytra is most strongly postero-dorsally translocated within the Trechiama Phyletic Series in a current sense (cf. Figs. 3d, g–h View Fig ). However, in Japanese trechines as a whole, the pore is most strongly postero-dorsally translocated in Suzuka S. Uéno, 1956 species, which belongs to the Trechoblemus Phyletic Series.

Geographic range. Northwestern area of Shikoku, southwestern Japan; northwest of the range of the nominate subgenus ( Fig. 7 View Fig ).

Etymology. This new subgenus is named in honour of Mr. Kôji Tanaka, who discovered the type species of this subgenus. Gender: masculine.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Carabidae

Genus

Nipponaphaenops

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