Stenus rorellus

Puthz, V., 2013, Übersicht über die orientalischen Arten der Gattung Stenus LATREILLE 1797 (Coleoptera, Staphylinidae) 330. Beitrag zur Kenntnis der Steninen, Linzer biologische Beiträge 45 (2), pp. 1279-1470 : 1307-1311

publication ID

https://doi.org/ 10.5281/zenodo.5300922

DOI

https://doi.org/10.5281/zenodo.10548221

persistent identifier

https://treatment.plazi.org/id/A92887EE-FFB1-700E-FF72-FC3F5E5070C2

treatment provided by

Marcus

scientific name

Stenus rorellus
status

 

Die S. rorellus View in CoL -Gruppe – Revision der Gruppe

Neue Definition: Kleine bis mittelgrosse (2,0- 4,8mm), makroptere Arten, Abdomen ungerandet, 4. Tarsenglied gelappt, Stirn meist sehr breit, 9. Sternum am Hinterrand gesägt, 10. Tergit am Hinterrand apikolateral spitz, median ausgerandet oder mit verschiedenen anker- und gabelförmigen Gestalten (Ausnahmen: S. pernix L. BENICK , S. scitulus L. BENICK ). Punktierung meist sehr dicht, mehrere Arten mit auffallend glänzender Beborstung. Fühler kurz, zurückgelegt nicht den Hinterrand des Pronotums erreichend. Stirn oft sehr breit. Pronotum wenig länger als breit.: Beine ohne besondere Sexualcharaktere, vordere Sternite ohne auffällige Merkmale, 8. Sternit meist nur flach ausgeradet. Aedoeagus meist mit spitzwinklig verengtem Medianlobus, Innensack breit sack-/taschenförmig, mit häkchenbesetzter Membran, stärker sklerotisierte Elemente meist fehlend, Parameren ohne besondere Merkmale.

Unter den Artengruppen mit ungerandetem Abdomen und gelapptem 4. Tarsenglied zeigt auch in der S. wasmanni View in CoL -Gruppe das 10. Tergit Besonderheiten (so S. spurius L. BENICK , S. spenceri PUTHZ, Abb. 48, 49), es ist aber hier apikolateral nur rundlich vorgezogen (nicht spitz), apikomedian ausgerandet, und der Aedoeagus ist anders gebaut, auch besitzen alle Arten dieser Gruppe eine schmälere Stirn.

Diese Artengruppe ist über das tropische Afrika, Madagaskar und die gesamte Orientalis verbeitet.

Key to the Oriental species of the group of Stenus rorellus FAUVEL View in CoL

(mDE = maximum distance between eyes)

1 Frons less broad (HW: mDE>1.80) (fig. 409)....................................................................2

- Frons broader (HW: mDE <1,80) (fig. 408)........................................................................9

2 Metallic species...................................................................................................................3

- Non-metallic species...........................................................................................................4

3 Larger, FB 2.5mm. Elytral punctation less dense, interstices in sutural third as wide or wider than diameter of punctures.: E (fig. 146). 4.2-5.0mm. Philippines (Mindanao)....... ......................................................................................................... S. viridis BERNHAUER View in CoL

- Smaller, FB 2.0mm. Elytral punctation denser, interstices in sutural third smaller than half diameter of punctures.: E (fig. 207). 4.2-5.0mm. Philippines (Luzon) ....................... ............................................................................................... S. smaragdinus BERNHAUER View in CoL

4 Punctation of frons sparse, interstices much wider than diameter of punctures. T 10 (fig. 148).: E (fig. 220). 3.8-4.5mm (FB 2.1mm). Philippines (Negros)............ sparsifrons n. sp

- Punctation of frons very dense, interstices smaller than half diameter of punctures. 3 very similar species.............................................................................................................5

5 Smaller, FB <1.7mm. T 10 (fig. 150).: E: fig. 222). 2.8-3.1mm (FB 1.6mm). Philippines (Luzon, Panay, Mindoro) ......................................... S. philippinus BERNHAUER View in CoL

- Larger, FB>2.0mm.............................................................................................................6

6: Apical portion of median lobe very narrow, blade-shaped in ventral view, curved dorsad in lateral view. T 10 (as in fig. 147). 3.8-4.1mm (FB 2.1mm). Malaysia (Sabah, Sarawak), Brunei, Indonesia (Kalimantan ....................................... S. denticaudatus PUTHZ View in CoL

-: Apical portion of median lobe narrowly lanceolate, straight in lateral view.................7

7: Apical excision of sternite 8 less deep (length of sternite: depth of emargination>2.8). T 10 (fig. 147). 3.8-4.6mm (FB 2.0-2.2mm. Philippines (Luzon, Samar, Mindoro, Antipolo, Siargo, Leyte, Dinagat, Mindanao), Indonesia (Sulawesi, Maluku (Halmahera)),?New Guinea (Papua). ............................................... S. simulans L. BENICK View in CoL

-: Apical excision of sternite 8 deeper (length of sternite: depth of emargination<2.5). T 10 (as in fig. 147).............................................................................................................8

8: Sternite 7 with a pubescent callus in proximal half of median impression. E: Apical portion of median lobe shorter (fig. 194). 4.0-4.7mm (FB 2.1-2.2mm. Philippines (Luzon)............................................................................................ S. tropicus BERNHAUER View in CoL

-: Median impression of Sternite 8 without a proximal callus. E: Apical portion of median lobe longer (fig. 195). 4.0-4.7mm (FB 2.1-2.2mm. Philippines (Negros, Palawan, Mindanao)......................................................................... S. wendeleri L. BENICK View in CoL

9 T 10 ancoriform (figs 149, 151-153).................................................................................10

- T 10 otherwise...................................................................................................................13

10 Larger, FB 1.8mm. T 10 (fig. 149).: E (fig. 5, ROUGEMONT 1986b). 3.2-3.6mm (FB 1.8-1.9mm). Nepal......................................................................... S. morvani ROUGEMONT View in CoL

- Smaller, FB <1.8mm.........................................................................................................11

11 Head broader (HW: EW>0.88)........................................................................................12

- Head less broad (HW: EW <0.88). T 10 (fig. 152).: Unknown. 3.4-4.1mm (FB 1.5- 1.6mm). Thailand........................................................................ S. ancorellus ROUGEMONT View in CoL

12 T 10 (fig. 151). 2.7-3.3mm (FB 1.5-1.6mm). Laos.............................. S. ancoratus View in CoL nov.sp.

- T 10 (fig. 154).: E (figs 205, 206). 2.8-3.2mm (FB 1.6-1.7mm). Indonesia (Lombok, Flores, Bali, Java)............................................................................... S. lombokensis PUTHZ View in CoL

13 Extremely densely punctate with remarkably shiny short golden pubescence. T 10 (fig. 153).: E (fig. 3, PUTHZ 1969a). 3.3-4.1mm (FB 1.9-2.0mm). N India...................... ............................................................................................................ S. furcillatus PUTHZ View in CoL - Densely or very densely punctate, pubescence less remarkable (except in S. auratilicomus nov.sp.).......................................................................................................14

14 T 10 (figs 156-166)...........................................................................................................15

- T 10 otherwise...................................................................................................................19

15 Smaller, FB <1.4mm. T 10 (fig. 156). 2.1-2.5mm (FB 1.3mm). Singapore........................... .............................................................................................. S. leontopolitanus CAMERON View in CoL

- Larger, FB 1.5mm...........................................................................................................16

16 T 10 (fig. 155),: E (fig. 8, PUTHZ 1971a). 3.2-4.1mm (FB 1.5-1.6mm). Sri Lanka, N India...................................................................................................... S. bihamatus PUTHZ View in CoL

- T 10 otherwise...................................................................................................................17

17 Apicolateral teeth of T 10 longer (figs 157, 158)..............................................................18

- Apicolateral teeth of T 10 shorter (figs 191-166).: E (fig. x). 2.7-4.0mm (FB 1.6- 1.9mm). Myanmar, India, Nepal, Thailand, Laos, Vietnam, Malaysia (mainland, Sabah, Sarawak), Indonesia (Sumatra).............................................. S. monomerus FAUVEL View in CoL

18 T 10 (fig. 158).: E (figs. 199, 200). 2.8-3.7mm (FB 1.5-1.8mm). Andaman Islands, Myanmar, Thailand, China (Hainan) .............................................. S. andamanensis PUTHZ View in CoL

- T 10 (fig. 157).: E (fig. 5, PUTHZ 1969a). 2.6-3.0mm (FB 1.5mm). N India, Nepal, Thailand, Laos. China (Hainan) ....................................................... S. sparsepilosus PUTHZ View in CoL

19 T 10 apicolaterally ± rounded, no distinct apicolateral teeth (figs 167-169). Large species...............................................................................................................................20

- T 10 with distinct apicolateral teeth. Mostly smaller species............................................22

20 Apical two fifths of femora infuscate, elytra punctation slightly less coarse. T 10 (fig. 168). 3.8-4.6mm (FB 2.0-2.2mm). Philippines (Luzon)........................ S. pernix L. BENICK View in CoL

- Femora very shortly infuscate apically, elytral punctation coarser. T 10 (figs 167, 169).............21

21 Punctation of pronotum very dense, slightly transversely coalescent. Head without metallic lustre. T 10 (fig. 169).: Unknown. 3.7-4.2mm (FB 2.2mm). Philippines (Luzon ................................................................................................. S. scitulus L. BENICK View in CoL

- Punctation of pronotum less dense, punctures distinctly delimited. Head with slight metallic lustre. T 10 (fig. 167).: Unknown. 3.7-4.3mm (FB 2.1mm). Philippines (Luzon)............................................................................................... S. aericeps L. BENICK View in CoL

22 Apicolateral teeth of T 10 very short, emagination between teeth less deep and nearly straight. T 10 (fig. 170).: E (fig. 198). 3.3-3.8mm (FB 1.7-1.8mm). Philippines (Luzon, Leyte, Negros, Mindanao) .................................................. S. ambiguus L. BENICK View in CoL

- Apicolateral teeth of T 10 longer, emargination between teeth deeper and more concave (figs 25-45)..........................................................................................................23

23 Lateral outline of posterior portion of T 10 slightly concave (figs 173-175)....................24

- Lateral outline of posterior portion of T 10 straight or convex (figs 176-191).................26

24 Larger, FB 1.7mm...........................................................................................................25

- Smaller, FB 1.7mm. Frons matt, interstices of punctation extremely narrow, frons broader (GW: mDE = 1.32). T 10 (fig. 174).: Unknown. 2.3-2.8mm (FB 1.4mm). China (Guizhou).................................................................................... S. hebetifrons View in CoL n. sp.

25 Head broader than elytra, elytra broader than long. T 10 (fig. 172).: E (fig. 202). 3.1-3.6mm (FB 1.7-1.8mm). Singapore, Indonesia (Sumatra) ............................................... ................................................................................................ S. fortepunctatus CAMERON View in CoL

- Head narrower than elytra, elytra longer than broad. T 10 (fig. 173.: E (fig. 215). 3.5-4.0mm (FB 1.8-2.0mm). Philippines (Luzon)...................................... S. schoedli View in CoL n. sp.

26 Apical teeth of T 10 shorter (figs 177-187........................................................................27

- Apical teeth of T 10 longer (figs 29, 188-191...................................................................37

27 Frons less broad: HW: mDE 1.60...................................................................................28

- Frons broader: HW: mDE <1.69.......................................................................................32

28 Smaller, FB 1.6-1.7mm. T 10 (figs 175, 184). Two very similar species..........................29 - Larger, FB 1.9mm...........................................................................................................30

29 In average smaller: 2.7-3.1mm (FB 1.5mm). Punctation of T 7 very dense, interstices much smaller than half diameter of punctures. T 10 (fig. 184).: E (fig. 203, 204). Japan, China (Shaanxi, Zhejiang), Korea................................................ S. confertus SHARP View in CoL

- In average larger: 2.9-3.5mm (FB 1.6-1.7mm). Punctation of T 7 less dense, shiny interstices can become as large as half diameter of punctures. T 10 (fig. 175).: E (fig. 221). Laos............................................................................................. S. kubani View in CoL n. sp.

30 Legs dark brown to black. T 10 (fig. 179). 3.5-4.5mm (FB 2.0mm). Laos, China (Yunnan) ................................................................................................... S. laoticus PUTHZ View in CoL

- Legs predominantly reddish yellow – brownish yellow, often with darker apices of femora...............................................................................................................................31

31 Frons broader (HW:mDE = 1.60, fig. 408), deeply concave, elytra shorter (EW: EL = 100). T 10 (fig. 180).: E (fig. 193). 4.0-4.7mm (FB 2.2-2.3mm). Philippines (Luzon) .... ............................................................................................. S. pseudotropicus BERNHAUER View in CoL

- Frons less broad (HW:mDE = 170, fig. 409), slightly concave, elytra longer (EW: EL = 0.96). T 10 (fig. 186).: E (fig. 13, PUTHZ 1998b). 3.3-4.1mm (FB 1.9mm). Malaysia ( Sarawak), Brunei...................................................................... S. pfeifferi PUTHZ View in CoL

32 Pubescence conspicuously golden, shiny. T 10 (fig. 181).: E (fig. 201). 2.9-3.5mm (FB 1.7mm). Myanmar.................................................................... S. aurantilicomus View in CoL n. sp.

- Pubescence not conspicuous, less shiny............................................................................33

33 Frons broader: HW: mDE <1.58.......................................................................................34

- Frons less broad: HW: mDE>1.60...................................................................................35

34 Smaller: FB 1.4-1.5mm. T 10 (fig. 177).: E (fig. 196). 2.2-2.7mm. Solomon Islands, New Caledonia, Papua New Guinea. .................................................. S. greensladei PUTHZ View in CoL

- Larger: FB 1.8mm...........................................................................................................36

35 Larger: FB 2.0mm. Frons deeply concave. T 10 (fig. 178).: Unknown. 3.7-4.3mm. N India, Myanmar.................................................................................... S. quisquilius PUTHZ View in CoL

- Smaller: FB 1.6-1.8mm. Frons broadly and shallowly elevated. T 10 (fig. 182.: E (fig. 5, PUTHZ 1994b). 2.9-3.3mm (FB1.6-1.8mm). Indonesia (Ceram)................................ ........................................................................................................ S. schillhammeri PUTHZ View in CoL

36 On average larger. T 10 (fig. 183).: E (figs 212, 213). 2.8-3.9mm (FB 1.5-2.0mm). Sri Lanka, India, Nepal, Myanmar, Thailand.......................... S. cribellatus MOTSCHULSKY View in CoL

- On average smaller. T 10 (fig. 185).: E (fig. 192). 2.5-3.0mm (FB 1.5mm). Sri Lanka................................................................................................ S. lacertoides NIETNER View in CoL

37 Extremely densely punctate with remarkably shiny short golden pubescence, interstices of punctures on tergite 7 smaller than half diameter of punctures, reticulate.......................................38

- Densely to very densely punctate, pubescence less remarkable, interstices on tergite 7 larger than half diameter of punctures, not reticulate........................................................39

38 Less large, FB 1.4-1.7mm. T 10 (fig 191), posterior margin between lateral teeth simple.: E (figs 43-45, 47, PUTHZ 1969b). 2.4-3.5mm. India, Pakistan, Nepal, Bhutan, Bangladesh, Myanmar, Thailand, Laos, Vietnam, China (Yunnan, Shanghai, Beijing), Malaysia (Sabah, Java, Lombok), Indonesia (Sulawesi, Timor, Lombok, Java), Philippines (Luzon, Mindoro), Australia, New Guinea, New Caledonia..................... .......................................................................................... S. rorellus View in CoL cursorius L. BENICK View in CoL

- Larger, FB 1.8-1.9mm. T 10 (fig. 189), posterior margin between lateral teeth serrate.: E (fig. 14, PUTHZ 1998b). 3.2-4.0mm. India (Goa) ............................... S. assingi PUTHZ View in CoL

39 Smaller: FB 1.3-1.5mm.....................................................................................................40

- Larger: FB 1.7-1.9mm.......................................................................................................41

40 Elytra larger (HW: EW <0,95). T 10 (fig. 171).: Unknown. 2.4-2.8mm (FB 1.5mm). Vietnam.................................................................................................... S. bicuspis PUTHZ View in CoL

- Elytra smaller (HW: EW>0,95). T 10 (fig. 176).: E (fig. 197). 2.3-2.8mm (FB 1.3- 1.5mm). Japan, China (Sichuan, Hainan, Taiwan, Hongkong, Zhejiang, Shanxi). ................ ............................................................................................................. S. dissimilis SHARP View in CoL

41 Punctation of frons regularly dense...................................................................................42

- Punctation of frons distinctly sparser medially (with a narrow shining area). T 10 (fig.190).: E (fig. 214). 3.4-4.0mm (FB 1.7-1.9mm). Philippines (Luzon, Virac, Samar, Mindoro, Mindanao, Leyte, Palawan), Malaysia ( Sarawak), Indonesia (Kalimantan, Sulawesi .......................................................................... S. nothus L. BENICK View in CoL

42 Apical teeth of T 10 longer (fig. 188), head broader (HW: EW>1.00).: E (fig. 4, PUTHZ 1981b). 3.0-3.5mm (FB 1.7mm). Sri Lanka, S India................. S. lewisiellus PUTHZ View in CoL

- Apical teeth of T 10 shorter (fig. 187), head less broad (HW: EW 0,.95).: E (fig. 22). 3.0-3.4mm (FB 1.6-1.8mm). Philippines (Luzon, Mindoro, Masbate, Negros, Palawan, Mindanao)......................................................................... S. sejugatus L. BENICK View in CoL

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Stenus

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