Trisunius, Assing, 2011

Assing, V., 2011, Trisunius gen. nov. from the southern East Palaearctic and the Oriental regions (Coleoptera: Staphylinidae: Paederinae: Medonina), Linzer biologische Beiträge 43 (1), pp. 195-220 : 218-219

Key to the species of Trisunius

1 Head glossy, without trace of microsculpture.....................................................................2

- Head with microsculpture...................................................................................................3

2 Body of more or less uniformly reddish-yellow coloration. Ƌ: sternite VII as in Fig. 77 ; posterior excision of sternite VIII small and narrow ( Fig. 78 ); ventral process of aedeagus slender and apically very acute both in lateral and in ventral view ( Figs 79- 80 ). Thailand............................................................................................. T. thaicus nov.sp.

- Head, pronotum, and abdomen dark-brown to blackish; elytra bicoloured, dark-brown to blackish-brown with the anterior and the posterior margins broadly reddish. Ƌ: sternite VII as in Fig. 54 ; posterior excision of sternite VIII broader and deeper ( Fig. 55 ); aedeagus of completely different shape ( Fig 56 ). China: Yunnan ................................. ..................................................................................................... T. appendiculatus nov.sp.

3 Species from northern India. Ƌ: sternite VII weakly modified ( Fig. 63 ); sternite VIII with moderately deep and almost V-shaped posterior excision ( Fig. 64 ); aedeagus small, approximately 0.3 mm long, and of distinctive shape ( Figs 65-66 ). India: Uttaranchal.................................................................................... T. monticola (CAMERON)

- Species from Yunnan, China. Ƌ: primary and secondary sexual characters different........4

4 Elytra usually conspicuously short ( Fig. 28 ), 0.70-0.75 times as long as pronotum and somewhat dilated posteriad (micropterous morph), rarely very long ( Fig. 29 ) and approximately 1.2 times as long as pronotum (macropterous morph). Relatively large (3.2-4.0 mm) and dark-coloured species; body usually brown to blackish-brown; elytra of uniform coloration. Ƌ: sternites VII and VIII as in Figs 30-31 ; aedeagus approximately 0.5 mm long and of distinctive shape ( Fig. 33 ) ........... T. discrepans nov.sp.

- Elytra in micropterous specimens at least approximately 0.8 times as long as pronotum and not distinctly dilated posteriad. On average smaller and often differently coloured species. Ƌ: secondary sexual characters different; aedeagus smaller and of different shape...................................................................................................................................5

5 Elytra usually bicoloured, dark-brown to blackish-brown with the posterior margin and/or the sutural portion reddish........................................................................................6

- Elytra of uniform coloration................................................................................................8

6 Elytra with the posterior sutural portion more or less distinctly reddish; posterior margin not - or only narrowly - reddish. Elytra long, 1.05-1.15 times as long as pronotum ( Fig. 35 ). Ƌ: sternite VII very weakly modified ( Fig. 37 ); posterior excision of sternite VII very small ( Fig. 38 ); ventral process of aedeagus apically very acute, slender in lateral view and spear-shaped in ventral view ( Figs 39-40 ).................................. .............................................................................................................. T. iaculatus nov.sp.

- Elytra with the posterior margin broadly reddish. Elytra shorter, 0.95-1.05 times as long as pronotum. Ƌ: primary and secondary sexual characters different..........................7

7 Head and pronotum reddish to reddish-brown. Ƌ: sternite VII very weakly modified ( Fig. 43 ); posterior excision of sternite VIII narrow and not very deep ( Fig. 44 ); aedeagus approximately 0.45 mm long and of distinctive shape ( Fig. 45 ). Recorded only from Bangma Shan ........................................................................ T. schuelkei nov.sp.

- Head and pronotum dark-brown to blackish-brown. Ƌ: sternite VII impressed in posterior median portion, on either side of this impression with extensive cluster of long setae, posterior margin concave in the middle ( Fig. 48 ); sternite VIII without pubescence along middle, posterior excision rather deep and V-shaped ( Fig. 49 ); aedeagus small, only approximately 0.3 mm long and of distinctive shape ( Figs 50-51 ) ..... .............................................................................................................. T. truncatus nov.sp. 8 Elytra usually short ( Fig. 18 ) and approximately 0.8 times as long as pronotum (micropterous morph), rarely very long ( Fig. 19 ) and approximately 1.1 times as long as pronotum (macropterous morph). Pronotum usually more or less distinctly microsculptured ( Fig. 21 ). Ƌ: sternite VII weakly modified ( Fig. 23 ); posterior excision of sternite VIII small ( Fig. 24 ); aedaegus of distinctive shape, ventral process laterally compressed ( Figs 25-26 ) ....................................................... T. cultellatus nov.sp.

- Elytra in submacropterous morph longer ( Figs 2 , 12 ), 0.8-0.9 mm as long as pronotum, and in macropterous morph shorter ( Fig. 11 ), 0.95-1.0 times as long as pronotum. Pronotum glossy, without microsculpture. Ƌ: sternite VII distinctly modified, with pronounced clusters of long setae ( Figs 6 , 13 ); posterior excision of sternite VIII deep and V-shaped ( Figs 7 , 14 ); ventral process of aedeagus of completely different shape, not compressed laterally......................................................................................................9

9 Ƌ: ventral process of aedeagus straight in lateral view and apically somewhat truncate in ventral view ( Figs 8-9 ) ................................................................... T. spathulatus nov.sp.

- Ƌ: ventral process of aedeagus sinuate in lateral view and apically rounded in ventral view ( Figs 15-16 ) ................................................................................... T. ligulatus nov.sp.

Figures

Figs 67-80: Trisunius thaicus nov.sp.: (67) habitus; (68) forebody; (69) median dorsal portion of head; (70) labrum; (71-72) mandibles; (73) maxilla; (74) labial palpi; (75) median dorsal portion of pronotum; (76) abdominal tergite VII; (77) male sternite VII; (78) male sternite VIII; (79-80) aedeagus in lateral and in ventral view. Scale bars: 67: 1.0 mm; 68: 0.5 mm; 69, 75-78: 0.2 mm; 70-74, 79-80: 0.1 mm.

Figs 50-56: Trisunius truncatus nov.sp. (50-51) and T. appendiculatus nov.sp. (52-56): (50, 56) aedeagus in lateral view; (51) aedeagus in ventral view; (52) habitus; (53) forebody; (54) male sternite VII; (55) male sternite VIII. Scale bars: 52: 1.0 mm; 53: 0.5 mm; 54-55: 0.2 mm; 50-51, 56: 0.1 mm. Ƌ: sternite VII in posterior median portion weakly impressed, on either side of this impression with moderately dense and not very evident cluster of long setae (Fig. 54); sternite VIII with not very deep, V-shaped excision (Fig. 55); aedeagus small, approximately 0.3 mm long; ventral process laterally compressed, of highly distinctive shape (Fig. 56).

Figs 57-66: Trisunius monticola (CAMERON): (57) habitus; (58) forebody; (59) median dorsal portion of head; (60-61) labium; (62) median dorsal portion of pronotum; (63) male sternite VII; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 57: 1.0 mm; 58: 0.5 mm; 59, 62-64: 0.2 mm; 60-61, 65-66: 0.1 mm.

Figs 27-33: Trisunius discrepans nov.sp.: (27) habitus (micropterous morph); (28) forebody (micropterous morph); (29) forebody (macropterous morph); (30) male sternite VII; (31) male sternite VIII; (32) male sternite IX; (33) aedeagus in lateral view. Scale bars: 27: 1.0 mm; 28-29: 0.5 mm; 30-32: 0.2 mm; 33: 0.1 mm.

Figs 34-40: Trisunius iaculatus nov.sp.: (34) habitus; (35) forebody; (36) labrum; (37) male sternite VII; (38) male sternite VIII; (39) aedeagus in lateral view; (40) ventral process of aedeagus in ventral view. Scale bars: 34: 1.0 mm; 35: 0.5 mm; 37-38: 0.2 mm; 36, 39-40: 0.1 mm.

Figs 41-49: Trisunius schuelkei nov.sp. (41-45) and T. truncatus nov.sp. (46-49): (41, 46) habitus; (42, 47) forebody; (43, 48) male sternite VII; (44, 49) male sternite VIII; (45) aedeagus in lateral view. Scale bars: 41, 46: 1.0 mm; 42, 47: 0.5 mm; 43-44, 48-49: 0.2 mm; 45: 0.1 mm.

Figs 17-26: Trisunius cultellatus nov.sp.: (17) habitus (micropterous morph); (18) forebody (micropterous morph); (19) forebody (macropterous morph); (20) median dorsal portion of head; (21) median dorsal portion of pronotum; (22) male sternite IX; (23) male sternite VII; (24) male sternite VIII; (25) aedeagus in lateral view; (26) ventral process of aedeagus in ventral view. Scale bars: 17: 1.0 mm; 18-19: 0.5 mm; 23-24: 0.2 mm; 20-22, 25-26: 0.1 mm.

Figs 1-9: Trisunius spathulatus nov.sp.: (1) habitus; (2) forebody; (3) median dorsal portion of head; (4) antenna; (5) labial palpi; (6) male sternite VII; (7) male sternite VIII; (8-9) aedeagus in lateral and in ventral view. Scale bars: 1: 1.0 mm; 2: 0.5 mm; 4, 6-7: 0.2 mm; 3, 5, 8-9: 0.1 mm.

Figs 10-16: Trisunius ligulatus nov.sp.: (10) habitus (macropterous morph); (11) forebody (macropterous morph); (12) forebody (submacropterous morph); (13) male sternite VII; (14) male sternite VIII; (15) aedeagus in lateral view; (16) ventral process of aedeagus in ventral view. Scale bars: 10: 1.0 mm; 11-12: 0.5 mm; 13-14: 0.2 mm; 15-16: 0.1 mm.