Coptocarpus microps, Will & Guéorguiev, 2021

Will, Kipling & Gueorguiev, Borislav, 2021, Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini), ZooKeys 1044, pp. 375-425 : 375

Coptocarpus microps sp. nov. Figs 22 , 32 , 49 , 53

Material examined.

Holotype: New Caledonia • ♂ " NEW CALEDONIA 21°10' Sx 165°18'E, Mt. Aoupinié, May-Oct 1992, R. Raven & E. Guilbert, rainforest, pitfalls " Holotype pinned, with genitalia in a separate microvial. Source collection QM, deposited MNHN. Type locality as given on label.

Diagnosis.

Similar to Adelopomorpha tethys in size and the lack of impressed striae 1-5, but with very small, flat eyes and male with protarsomeres 1-3 expanded. Both A. glabra and A. tuberculata also lack impressed striae, but are much smaller, and in C. microps the male has a pair of setae on ventrite 6 that is lacking in those species. The aedeagus is decisively different from all other species.

Description.

Habitus. Small sized, BL: 6.80 mm and BW: 3.00 mm, ovate, convex body. Color and luster. Head, prothorax, and legs deep reddish brown. Pronotal disc and apical portion of prosternal process darker, nearly black. Elytra and ventrites mostly black, paler reddish black on elytral epipleura and near humeri, and medially on ventrite 6; antennae and palpi reddish brown. Integument moderately dull, without spectral iridescence. Microsculpture and punctation. Dorsal surface of head and pronotum with prominent isodiametric meshes; elytra with somewhat irregular and slightly less prominent isodiametric meshes throughout, more irregular or very slightly stretched in apical 1/3; ventral surface with scarcely-visible sculpticells or mostly sculpticells not apparent. Head lacking macropunctation or wrinkles on vertex; head, pronotum and elytra with scattered micropunctation, only on elytra do micropunctures and sculpticells form some irregular rosettes (sensu Spence 1983); abdominal ventrites 1-3 at sides with several large punctures (more irregular in ventrite 3), ventrites 4 and 5 at sides with few coarse wrinkles, and ventrite 6 nearly smooth. Chaetotaxy. Labrum with six setae of nearly equal length each in its own socket, lateral setae slightly larger than medial four. Clypeal setae present near apicolateral corners. Single supraorbital seta present over each eye. Elytron without discal setiferous punctures. Abdominal ventrites 1-5 without ambulatory setae. Head. Approximately 1/3 as wide as pronotum (Suppl. material 2: Table S1). Eye very small, slightly protruded, EyW/HW: 1.17. Labrum anterior margin straight. Frontoclypeal sutures not evident, marked by a very broad, shallow depression. Antenna moderately long, with last segment exceeding pronotal base and pubescence starting from antennomere 4. Last labial palpomere slightly swollen, blunt at apex, slightly longer than penultimate one. Mentum without paramedial border. Thorax. Pronotum just slightly less than 1 2/3 wider than long (PW/PL: 1.61); width at apex 2 1/2 × less than at widest point (PW/PA: 2.52). Disc with middle line fine, very shallow, and ended well before apical and basal margins, without apical transverse impression; anterior angles not prominent, scarcely convex at tips, anterior margin shallowly concave, submarginal sulcus present near angles, lacking in middle 1/3. Prosternal process rounded, bordered throughout. Mesosternum with single, low medial tubercle. Metepisternum 1 1/3 × wider than long, with lateral margin nearly straight, coadunation with epipleuron along entire length. Elytra. Approximately 1/3 longer than wide (EL/EW: 1.27). Basal margin forming extremely minute, blunt denticle at shoulder, ended medially at level of parascutellar punctures, joined to parascutellar puncture fovea by a short striole. Humeral submarginal carina absent. Apical sinuation evident but shallow. Disc with no elytral striae impressed, those marked by very shallow, fine punctures; parascutellar striole absent. Granulation in marginal furrow discontinuous, interrupted broadly at midlength. Legs. Mesotibia notably dilated apically. Basomesotarsus and basometatarsus glabrous dorsally; basomesotarsus broadly flattened, with seven or eight very short, stout setae ventrally. Female genitalia. Female specimens unknown. Male genitalia. Median lobe of aedeagus in lateral view long, almost straight, with thin apex (Fig. 32 ); lobe in dorsal view, with apical lamella well bent to right, with right side concave and left side straight (Fig. 22 ); basal bulb rounded dorsally; sclerotized portion of endophallus visible in repose, with three sclerites, two near base of endophallus and one near apex.

Etymology.

The specific epithet Coptocarpus microps is Latin for small eyes and draws attention to the extraordinary small eyes in these beetles (Fig. 49 ). It is treated as an adjective in the nominative singular.

Figures

Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima.

Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus.

Figures 49 - 51. Head in left lateral view of 49 Coptocarpus microps sp. nov. 50 C. cyllodinus (Fauvel) 51 C. magnus sp. nov. Arrow indicates location of notch beneath the eye.

Figures 52, 53. Maps of localities of New Caledonian Oodini.