Evanirvana Hill, 1973

Gonçalves, Clayton C., Dietrich, Christopher H. & Takiya, Daniela M., 2019, Contribution to the tribe Pentoffiini (Cicadellidae: Evacanthinae) with descriptions of two new species and the male of Evanirvana aurea Hill, 1973, Zootaxa 4688 (2), pp. 271-281 : 272-273

Evanirvana Hill, 1973

Type-species Evanirvana aurea Hill, 1973 by monotype

Diagnosis. Medium-sized leafhoppers. Crown ( Figs 35, 39 ) with median longitudinal carina complete extending from the anterior to posterior margin. Pronotum and base of clavus ( Figs 35, 36 ) punctate. Forewing ( Figs 2 , 16 ) costal margin with supranumerary R branches along apical half. Subgenital plate ( Figs 4 , 18 ) narrow and elongated. Aedeagus ( Figs 8 , 22 ) with elongate dorsally curved pair of atrial processes; shaft with apical portion expanded with pair of lateral processes.

Coloration. Dull yellow ( Figs 11–13 , 33–40 ) with dark maculae on crown that may extend to pronotum ( Figs 35, 39 ). Forewing milky hyaline; male with smoky median longitudinal band extending from the clavus to apical cells, absent in female ( Figs 2 , 11 , 16 ).

External morphology. Head ( Figs 35, 39 ), in dorsal view, distinctly narrower than pronotum. Crown ( Fig. 35, 39 ) moderately produced anteriorly, expanded laterad anterad of eyes; coronal suture carinate and elevated towards apex; surface depressed between coronal suture and lateral margins of crown, texture with numerous longitudinal and oblique striations; anterior margin rounded or angled with distinct carina at crown-face transition; lateral margins, adjacent to eyes, raised and carinated. Frons, in lateral view ( Figs 36, 40 ), oblique and slightly convex; inflated; with median longitudinal carina dorsally. Antenna moderately long; flagellum approximately one-fourth length of body. Antennal ledge carinate, in frontal view ( Figs 1 , 13 , 15 ) narrow, flap-like; in lateral view ( Figs 36, 40 ) slightly oblique. Gena broadly rounded, not emarginated below eye ( Figs 1 , 13 , 15 ). Clypeus ( Figs 1 , 13 , 15 ) rectangular; with parallel lateral margins; ventral margin straight or slightly arched.

Pronotum ( Figs 35, 39 ) punctate; lateral margins carinate and strongly convergent anteriorly. Forewing ( Figs 2 , 16 ) hyaline; venation distinct; costal margin with supranumerary R branches along apical half; s crossvein slightly anterior to r-m 2; r-m 1 absent; three or more m-cu crossveins present; three anteapical cells (median one open or closed) and four apical cells; inner apical cell narrow and elongate; clavus with base punctate; appendix narrow, extending to second apical cell. Hind wing with veins R 4+5 and M 1+2 separated by short crossvein; crossvein m-cu short. Hind femur with macrosetal formula 2+1+1; tibia with AD row with two or three intercalary microsetae between macrosetae; tibia with AV row with 5–6 macrosetae restricted to distal half; first tarsomere with two parallel rows of small setae on plantar surface; apex with three platellae; second tarsomere apex with two apical platellae.

Male terminalia. Male pygofer ( Figs 3 , 17 ) slightly angulate posterodorsally; without lobes or processes; macrosetae restricted to apical area. Valve subrectangular; with sclerotized median line. Subgenital plate ( Figs 3–4 , 17–18 ) narrow; ventral surface with scattered fine setae. Connective ( Figs 5 , 19 ) T-shaped. Style ( Figs 5–6 , 19–20 ) with apodeme long; apophysis short and stout; apex hooked on inner side and with distinct long spine on outer side. Aedeagus ( Figs 7–8 , 21–22 ) with pair of long dorsally curved atrial processes; shaft depressed dorsoventrally, with pair of preapical lateral processes; gonopore apical.

Female terminalia. Sternite VII ( Fig. 9 ) wider than long; posterolateral angles reduced. First valvula ( Fig. 10 ) sculpturing pattern with dorsal area strigate and ventroapical area maculose. Second valvulae ( Hill, 1973: Figs C, D) with irregular teeth crowded apically.

Distribution. Brazil: Rio de Janeiro, Santa Catarina; Peru: Madre de Dios.

Figures

FIGURES 33−44. 33−36, Evanirvana aurea Hill, 1973, male. 33, dorsal habitus. 34, lateral habitus. 35, head and prothorax, dorsal view. 36, head and prothorax, lateral view. 37−40, Evanirvana tambopata sp. nov., male holotype. 37, dorsal habitus. 38, lateral habitus. 39, head and prothorax, dorsal view. 40, head and prothorax, lateral view. 41−44, Pentoffia santosi sp. nov., male holotype. 41, dorsal habitus. 42, lateral habitus. 43, head and prothorax, dorsal view. 44, head and prothorax, lateral view.

FIGURES 1−8. Evanirvana aurea Hill, 1973, male. 1, head, ventral view. 2, forewing. 3, pygofer, valve, subgenital plate, and anal tube, lateral view. 4, subgenital plate, ventral view. 5, connective and styles, dorsal view. 6, style, lateral view. 7, aedeagus, lateral view. 8, aedeagus, caudal view.

FIGURES 15−22. Evanirvana tambopata sp. nov., male holotype. 15, head, ventral view. 16, forewing. 17, pygofer, valve, and subgenital plate, lateral view. 18, subgenital plate, ventral view. 19, connective and styles, dorsal view. 20, style, lateral view. 21, aedeagus, lateral view. 22, aedeagus, caudal view.

FIGURES 9−14. Evanirvana aurea Hill, 1973, holotype female (USNM 01513539). 9, sternite VIII, ventral view. 10, pygofer and first valvulae, lateral view. 11, habitus, dorsal view. 12, habitus, lateral view. 13, head, ventral view. 14, specimen labels.