Cancilla planicostata (Bellardi, 1887)

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 54

publication ID	https://doi.org/10.11646/zootaxa.4983.1.1
publication LSID	lsid:zoobank.org:pub:6A4778D6-195A-4AB1-AA1E-7D8000185B28
DOI	https://doi.org/10.5281/zenodo.5044092
persistent identifier	https://treatment.plazi.org/id/A82A87E9-8A23-387B-FF4D-FF65FD35FF31
treatment provided by	Plazi (2021-06-10 00:51:57, last updated 2024-11-24 21:02:59)
scientific name	Cancilla planicostata (Bellardi, 1887)
status

Treatment

Cancilla planicostata (Bellardi, 1887) View in CoL

Figs 14E View FIGURE 14 1 –E View FIGURE 1 2 View FIGURE 2 , F 1 –F View FIGURE 1 2 View FIGURE 2 , G 1 –G View FIGURE 1 2 View FIGURE 2 , H, 7I View FIGURE 7 1 –I View FIGURE 1 2 View FIGURE 2 , J 1 –J View FIGURE 1 2 View FIGURE 2

Mitra (Nebularia) scrobiculata Brocc. — Hoernes & Auinger 1880: 80, pl. 9, figs 17a–b [non Brocchi, 1814].

Mitra exarata Bell. — Bellardi 1887a: 84, pl. 4, fig. 53 [non A. Adams, 1853].

* Mitra planicostata Bell. View in CoL — Bellardi 1887b: 5, pl. 2, fig. 9.

Mitra (Mitra) exarata Bell. — Boettger 1906: 8 [non A. Adams, 1853].

M [itra]. (T [iara].) planicostata Blld. View in CoL — Sieber 1958a: 154.

M [itra]. (T [iara].) grateloupi Orb. — Sieber 1958a: 154 [non d’Orbigny, 1852].

Mitra (Tiara) grateloupi Orb. — Sieber 1958b: 149 [non d’Orbigny, 1852].

Mitra (Tiara) grateloupi d’Orbigny, 1852 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 157, pl. 42, fig. 1.

Mitra exarata Bellardi, 1887 — Ferrero Mortara et al. 1981: 162, pl. 50, figs 1a–b.

Mitra planicostata Bellardi, 1887 View in CoL — Ferrero Mortara et al. 1981: 162, pl. 50, figs 2a–b.

Cancilla planicostata Bellardi, 1887 View in CoL — Cavallo & Repetto, 1992: 118, fig. 303.

Mitraria (Tiara) grateloupi d’Orbigny — Schultz 1998: 70, pl. 28, fig. 9.

Cancilla planicostata (Bellardi, 1887) View in CoL — Chirli 2002: 45, pl. 22, figs 10–11.

Cancilla planicostata (Bellardi, 1887) View in CoL — Landau et al. 2013: 211, pl. 33, figs 4–8, pl. 80, fig. 7.

Type material. Syntype ( BS.019.01.190) illustrated in Ferrero Mortara et al. (1981, pl. 50, figs 2a–b), Pliocene , Vezza Alba , Italy, stored in the Museo Regionale di Scienze Naturali, Torino .

Additional material. NHMW 1854/0035/0092, SL: 45.7 mm, MD: 11.9 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 17a–b), figs 14F 1 –F 2; NHMW 2020/0112/0001, SL: 35.5 mm, MD: 8.5 mm, Coşteiu de Sus ( Romania), figs 14G 1 –G 2, figs 7I 1 –I 2; NHMW 2020/0112/0002, SL: 35.6 mm, MD: 9.2 mm, Coşteiu de Sus ( Romania), figs 14E 1 –E 2; NHMW 2020/0112/0003, SL: 40.3 mm, MD: 10.9 mm, Coşteiu de Sus ( Romania), fig. 14H; NHMW 2020/0112/0002, SL: 37.5 mm, MD: 9.9 mm, Coşteiu de Sus ( Romania), figs 7J 1 –J 2.

Revised description (of Paratethyan material). Shell moderately large, slender fusiform with narrowly incised suture. Protoconch high conical of about three weakly convex whorls. Teleoconch of eight whorls; spire whorls nearly straight sided with periphery at abapical suture; rarely slightly convex. First teleoconch whorl weakly cancellate with predominant spiral sculpture composed of five spiral cords. On second teleoconch whorl sixth spiral cord appears at abapical suture, intermediate whorls with six fully exposed cords, seventh cord appears on penultimate whorl. Spiral cords broad, with flat tops and rounded edges, separated by narrow, punctate spiral grooves. Last whorl elongate, weakly convex, slowly contracting with shallow concavity at base. About 18 to 20 spiral cords on last whorl. Outer lip thin, weakly crenulated by terminations of spiral grooves. Aperture moderately narrow. Columellar callus narrow, thin, sharply delimited, prominent fasciole. Columella with three oblique folds, weakening abapically. Siphonal canal long, moderately narrow, distinctly twisted, with deep siphonal notch. Colour pattern in UV light consisting of dark, broad, weakly undulating axial stripes or flammulae separated by light interspaces. Less intense dark spiral stripes in spiral grooves.

Shell measurements and ratios. SL = 31.1–45.7 mm, MD: 8.5–11.9 mm; AA = 24–28°, SL/ MD: 3.6–4.2, AL/AW: 5.4–6.5, AH/S: 2.0–2.1.

Discussion. The species displays some variability in its spiral sculpture. The adapical two spiral cords may amalgamate and the spiral grooves on the last whorl may become subobsolete along the periphery. The overall shape and sculpture suggest a relationship with the Cancilla scrobiculata group [e.g. C. praescrobiculata ( Toldo, 1889) , C. scrobiculata ( Brocchi, 1814) , C. pulcherrima (Bellardi, 1887) ]. It is clearly separated from these species by its much smaller size and the less elongate outline. The axial colour pattern of Cancilla planicostata , was also documented by Landau et al. (2013) from Serravallian specimens from Turkey, supporting our assumption that both occurrences are conspecific.

Palaeoenvironment. The specimens from the Vienna Basin were collected from basinal clays of the Baden Formation , indicating middle to outer neritic environments with up to 250 m water depth ( Hohenegger et al. 2008) .

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau ( Schultz 1998), Făget Basin: Coşteiu de Sus, Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1880); Dacian Basin: Staropatica ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Serravallian (middle Miocene): Karaman Basin: Akboðazi, Akpýnar, Lale, Pýnarlar Yaylasý, Seyithasan, Tilkikaya ( Turkey) ( Landau et al. 2013); Tortonian (late Miocene): Po Basin: Montegibbio ( Italy) ( Davoli 2000; Zanclean (Pliocene): Roussillon Basin, France ( Chirli & Richard 2008); central Mediterranean, Italy ( Bellardi 1887b; Sacco 1904; Chirli 2002).

References

Bellardi, L. (1887 a) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 5; Mitridae. Ermanno Loescher, Torino, 85 pp. [https: // www. biodiversitylibrary. org / item / 45379 # page / 9 / mode / 1 up]

Bellardi, L. (1887 b) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 5; Mitridae (continuazione). Ermanno Loescher, Torino, 85 pp. [https: // www. biodiversitylibrary. org / item / 45379 # page / 113 / mode / 1 up]

Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]

Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Stamperia Reale, Milano, 2 + 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569

Cavallo, O. & Repetto, G. (1992) Conchiglie fossili del Roero. Atlante iconografico. Associazione Naturalistica Piemontese Memorie (Associazione Amici del Museo ' Federico Eusebio'), 2, 1 - 251.

Chirli, C. (2002) Malacofauna Pliocenica Toscana. Vol. 3. Superfamiglia Muricoidea-Cancellarioidea. C. Chirli, Firenze, 92 pp.

Chirli, C. & Richard, C. (2008) Les Mollusques plaisanciens de la Cote d'Azur. C. Chirli, Tavernelle, 128 pp.

d'Orbigny, A. (1852) Prodrome de Paleontologie Stratigraphique Universelle des Animaux Mollusques & Rayonnes faisant suite au Cours Elementaire de Paleontologie et de Geologie Stratigraphique. Vol. 3. Victor Masson, Paris, 189 pp. [https: // www. biodiversitylibrary. org / item / 98989 # page / 9 / mode / 1 up]

Davoli, F. (2000) I gasteropodi mitriformi del Tortoniano di Montegibbio (Subappennino modenese). Bolletino della Societa Paleontologica Italiana, 39 / 2, 165 - 215.

Ferrero Mortara, E., Montefameglio, L., Pavia, G. & Tampieri, R. (1981) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte I. Museo Regionale di Scienze Naturali, Catalogo, 6, 1 - 327. [https: // www. biodiversitylibrary. org / item / 269998 # page / 5 / mode / 1 up]

Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405

Hohenegger, J., Andersen, N., Baldi, K., Coric, S., Pervesler, P., Rupp, C. & Wagreich, M. (2008) Palaeoenvironment of the Early Badenian Middle Miocene in the southern Vienna Basin Austria - multivariate analysis of the Baden-Sooss section. Geologica Carpathica, 59, 461 - 487. [http: // www. geologicacarpathica. com / browse-journal / volumes / 59 - 5 / article- 458 /]

Kojumdgieva, E. M. & Strachimirov, B. (1960) Les fossiles de Bulgarie. VII. Tortonien. Academie des Sciences de Bulgarie, Sofia, 317 pp.

Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576

Sacco, F. (1904) I molluschi dei terreni terziari del Piemonte e della Liguria. Parte 30. Aggiunte e correzioni (con 1400 figure). Considerazioni generali. Indice generale dell'opera. C. Clausen, Torino, 203 + xxxvi pp., 31 pls. [https: // www. biodiversitylibrary. org / item / 46270 # page / 90 / mode / 1 up]

Schultz, O. (1998) Tertiarfossilien Osterreichs, Wirbellose, niedere Wirbeltiere und marine Saugetiere; schone, interessante, haufige und wichtige Makrofossilien aus den Bestanden des Naturhistorischen Museums Wien und Privatsammlungen; eine Bilddokumentation. Goldschneck-Verlag, Weinstadt, 159 pp.

Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Sieber, R. (1958 b) Die Tortonfauna von Steinabrunn bei Drasenhofen (Bez. Mistelbach, N. - O.). Verhandlungen der Geologischen Bundesanstalt, 1958, 142 - 155. [https: // opac. geologie. ac. at / ais 312 / dokumente / VH 1958 _ 142 _ A. pdf]

Toldo, G. (1889) Mitridae del Miocene superiore di Montegibbio. Bollettino della Societa Malacologica Italiana, 14, 144 - 150. [https: // www. biodiversitylibrary. org / item / 121430 # page / 148 / mode / 1 up]

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 7. Colour patterns in UV light. A1–A2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0002. B1–B2. Domiporta amoena nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0001. C1–C2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 1868/0001/0403. D1–D2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0003. E1–E2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0002. F1–F2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0003. G1–G2. Domiporta pulchra nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0001. H1–H2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0004. I1–I2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0001. J1–J2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0002. K1–K2. Domiporta turpis nov. sp., holotype, Baden (Austria), NHMW 1846/0037/0098ª. L1–L2. Episcomitra pseudoincognita nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0122/0001.

FIGURE 14. A1–A2. Cancilla exornata (Bellardi, 1887), Baden (Austria), NHMW 2020/0111/0003. B1–B2. Cancilla exornata (Bellardi, 1887), Baden (Austria), NHMW 2020/0111/0001. C1–C2. Cancilla exornata (Bellardi, 1887), Baden (Austria), NHMW 2020/0111/0004. D. Cancilla exornata (Bellardi, 1887), Baden (Austria), NHMW 2020/0111/0002. E1–E2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus(Romania), NHMW 2020/0112/0002.F1–F2. Cancilla planicostata (Bellardi, 1887), Lăpugiu de Sus (Romania), NHMW 1854/0035/0092. G1–G2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Roma- nia), NHMW 2020/0112/0001. H. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0003. I1–I2. Cancilla praescrobiculata (Toldo, 1889), lectotype of Mitra orientalis Oppenheim, 1918 [non Griffith & Pidgeon, 1834], Baden (Austria), NHMW 1846/0037/0098. J1–J2. Cancilla praescrobiculata (Toldo, 1889), Baden-Sooss (Austria), NHMW 2020/0114/0001. K1–K. Cancilla praescrobiculata (Toldo, 1889), Baden-Sooss (Austria), NHMW 2020/0114/0002. L. Cancilla praescrobiculata (Toldo, 1889), Baden-Sooss (Austria), NHMW 2020/0114/0003.