Calcimitra bellardii ( Hoernes & Auinger, 1880 )

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 25-27

publication ID	https://doi.org/10.11646/zootaxa.4983.1.1
publication LSID	lsid:zoobank.org:pub:6A4778D6-195A-4AB1-AA1E-7D8000185B28
DOI	https://doi.org/10.5281/zenodo.5044050
persistent identifier	https://treatment.plazi.org/id/A82A87E9-8A0E-3857-FF4D-FF65FC1EFE82
treatment provided by	Plazi (2021-06-10 00:51:57, last updated 2024-11-24 21:02:59)
scientific name	Calcimitra bellardii ( Hoernes & Auinger, 1880 )
status

Treatment

Calcimitra bellardii ( Hoernes & Auinger, 1880) View in CoL

Figs 6A View FIGURE 6 1 –A View FIGURE 1 2 View FIGURE 2 , B 1 –B View FIGURE 1 2 View FIGURE 2 , C 1 –C View FIGURE 1 2 View FIGURE 2 , D 1 –D View FIGURE 1 2 View FIGURE 2 , E 1 –E View FIGURE 1 2 View FIGURE 2 , F 1 –F View FIGURE 1 2 View FIGURE 2

[ Mitra View in CoL ] subulata Partsch— Hörnes 1848: 17 [nomen nudum] [non Lamarck, 1811].

Mitra Bronni Micht. View in CoL — Hörnes 1852b: 103, pl. 10, figs 22–24 [non Michelotti, 1847].

[ Mitra View in CoL ] Bronni Micht. — Auinger 1871: 8 [non Michelotti, 1847].

[ Mitra View in CoL ] Bellardii nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Bellardii View in CoL nov. form.— Hoernes & Auinger 1880: 78 (pars) [non pl. 9, figs 15–16 = Episcomitra antibellardii View in CoL nov. sp.].

Mitra bellardii Hö. Au. View in CoL — Boettger, 1902: 13.

Mitra (Mitra) subfusulus n. nom.— Boettger 1906: 5, nov. nom. pro Mitra bellardii Hoernes & Auinger, 1880 View in CoL .

M [itraria]. (M [itraria].) bellardii View in CoL (R. Hörn, et Au.)— Sieber 1958a: 153.

Mitra (Mitraria) auingeri View in CoL nom. nov. —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 159, pl. 42, fig. 7 [nov. nom. pro Mitra bellardii Hoernes & Auinger, 1880 View in CoL ].

Mitra bellardii Hoernes et Auinger View in CoL — Csepreghy-Meznerics 1969: 92, pl. 5, fig. 31.

Mitra bellardii Hoernes et Auinger View in CoL — Csepreghy-Meznerics 1972: 31, pl. 14, fig. 14.

Cancilla bellardii ( Hoernes & Auinger, 1880) — Cernohorsky 1991: 36.

non Mitra (Mitra) bellardii Hoernes et Auinger, 1880 View in CoL — Glibert 1952a: 118, pl. 9, fig. 4.

non Mitra (Nebularia) scrobiculata var. bellardii Hoernes & Auinger, 1880 View in CoL — Strausz 1966: 365, pl. 26, figs 9–10 [= Cancilla nanostriatula View in CoL nov. sp.].

Type material. Lectotype (designated herein): NHMW 2020 View Materials /0135/0001, SL: 24.4 mm, MD: 6.8 mm, Baden ( Austria), illustrated in Hörnes (1852b, pl. 10, fig. 23), figs 6F 1 –F 2 . Paralectotype: NHMW 1846 View Materials /0037/0107a, SL: 23.7 mm , MD: 6.3 mm, Baden ( Austria), illustrated in Hörnes (1852b, pl. 10, fig. 24), figs 6E 1 –E 2 .

Additional material. SFM 12.2243a, SL: 21.8 mm, MD: 5.5 mm, Senckenberg Museum , Frankfurt / Main, Germany , Coşteiu de Sus ( Romania), figs 7A 1 –A 2; SFM 12.2243a, SL: 19.9 mm, MD: 6.0 mm, Senckenberg Museum , Frankfurt / Main, Germany , Coşteiu de Sus ( Romania), figs 6B 1 –B 2; SFM 12.2243a, SL: 18.7 mm, MD: 5.7 mm, Senckenberg Museum , Frankfurt / Main, Germany , Coşteiu de Sus ( Romania), figs 6C 1 –C 2; NHMW 2020 View Materials /0133/0001, SL: 23.4 mm, MD: 6.4 mm, Lăpugiu de Sus ( Romania), figs 6D 1 –D 2; NHMW 2020 View Materials /0133/0002, SL: 20.1 mm, MD: 5.8 mm, Lăpugiu de Sus ( Romania); NHMW 2020 View Materials /0133/0003, SL: 21.9 mm, MD: 6.5 mm, Lăpugiu de Sus (Romania); NHMW 1870 View Materials /0033/0033, 44 specimens, Lăpugiu de Sus ( Romania) .

Description. Shell glossy, small, slender fusiform. Protoconch high conical of three smooth convex whorls. Teleoconch of seven whorls. First teleoconch whorls subcylindrical, slightly gradate, weakly canaliculate suture. Later spire whorls straight sided to weakly convex, with periphery at abapical suture. About eight prominent, convex axial ribs on first teleoconch whorl separated by slightly wider interspaces. Axial ribs weakening on third teleoconch whorl crossed by three prominent, slightly punctate spiral grooves placed on adapical half of whorl. Additional weaker spiral grooves on penultimate whorl. Last whorl nearly straight sided above periphery, slowly contracting into long base; spiral sculpture subobsolete along periphery. Numerous low, convex spiral cords on base and fasciole. Interspaces slightly punctate with delicate axial growth lines. Aperture elongate, narrow to moderately narrow, posteriorly angulated with indistinct anal notch. Columellar callus thin, broad, glossy. Columella with three weak, oblique columellar folds. Outer lip thin. Siphonal canal long, bent to the left, strongly twisted with moderately deep siphonal notch.

Shell measurements and ratios. SL: 18.7–25.5 mm, MD: 5.5–7.6 mm, AA = 24–27°, SL/MD: 3.5–3.8, AL/ AW: 4.8–5.9, AH/S: 1.9–2.2.

Discussion. Hoernes & Auinger (1880) referred to the specimens illustrated by Hörnes (1852, pl. 10, figs 22–24) when establishing Mitra bellardii as new name [“ die von Baden stammenden Originale ” (the original specimens from Baden)]. Therefore, these specimens are syntypes. Herein, we select the specimen illustrated in Hörnes (1852b, pl. 10, fig. 23) as lectotype and specimen on fig. 24 as paralectotype. The third syntype ( Hörnes 1852b, pl. 10, fig. 22) is lost. Unfortunately, Hoernes & Auinger (1880) illustrated two specimens from Lăpugiu de Sus (Romania) as Mitra bellardii , stating that these specimens differ in their large size ( Hoernes & Auinger 1880, pl. 9, figs 15–16). The Lăpugiu de Sus specimens are not conspecific with Calcimitra bellardii , but influenced the concept of ‘ Mitra bellardii ’ of subsequent authors (e.g. Glibert 1952; Janssen 1972). These two specimens are described herein as Episcomitra antibellardii nov. sp.

Boettger (1906) assumed that Mitra bellardii Hoernes & Auinger, 1880 was preoccupied by Foresti (1879) and introduced Mitra subfusulus as new name. Foresti (1879: 115), however, described his species as Mitra bellardiana and therefore, there is no homonymy. For the same reason, Mitra (Mitraria) auingeri Kojumdgieva in Kojumdgieva & Strachimirov 1960 is a superfluous replacement name. Calcimitra fusulus ( Cocconi, 1873) , from the Mediterranean Pliocene, differs only in its larger size (SL = 40 mm) and the less twisted siphonal canal (see Cavallo & Repetto 1992, fig. 301; Chirli 2002, pl. 22, figs 5–9).

Calcimitra bellardii View in CoL differs from all the species in Bellardi’s (1887b: 12) “ 46 a Serie ” (see generic discussion) in having the spirals obsolete, subobsolete, or greatly reduced in number.

Palaeoenvironment. The clay of the Baden Formation formed in middle to outer neritic settings with up to 250 m water depth ( Hohenegger et al. 2008). Extant Calcimitra species are deep water species ( Fedosov et al. 2018).

Distribution in Central Paratethys. Badenian (middle Miocene): North-Alpine-Carpathian Foredeep: Jerutek at Lysice, Hrušovany nad Jevišovkou, Jaroměřice nad Rokytnou ( Czech Republic) ( Hoernes & Auinger 1880); Vienna Basin: Baden, Gainfarn, Niederleis ( Austria) ( Hoernes & Auinger 1880); Eisenstadt-Sopron Basin: Forchtenau ( Austria) ( Hoernes & Auinger 1880); Bükk Mountains ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Hoernes & Auinger 1880); Dacian Basin: Staropatica, Yasen, Urovene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

References

Auinger, M. (1871) Tabellarisches Verzeichnis der bisher aus den Tertiarbildungen der Markgrafschaft Mahren bekannt gewordenen fossilen Conchylien auf Grundlage der Sammlung des k. k. Hof-Mineralien-Cabinetes. Naturforschender Verein Brunn, Verhandlungen, 9, 1 - 27. [https: // opac. geologie. ac. at / ais 312 / dokumente / Auinger _ 1871 _ Tertiaerbildungen. pdf]

Bellardi, L. (1887 b) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 5; Mitridae (continuazione). Ermanno Loescher, Torino, 85 pp. [https: // www. biodiversitylibrary. org / item / 45379 # page / 113 / mode / 1 up]

Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]

Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]

Cavallo, O. & Repetto, G. (1992) Conchiglie fossili del Roero. Atlante iconografico. Associazione Naturalistica Piemontese Memorie (Associazione Amici del Museo ' Federico Eusebio'), 2, 1 - 251.

Cernohorsky, W. O. (1991) The Mitridae of the World. Part 2. The Subfamily Mitrinae concluded and Subfamilies Imbricariinae and Cylindromitrinae. Monographs of Marine Mollusca 4, 1 - 164.

Chirli, C. (2002) Malacofauna Pliocenica Toscana. Vol. 3. Superfamiglia Muricoidea-Cancellarioidea. C. Chirli, Firenze, 92 pp.

Cocconi, G. (1873) Enumerazione sistematica dei Molluschi Miocenici e Pliocenici delle provincie di Parma e di Piacenza. Gamberini e Parmeggiani, Bologna, 368 pp. [https: // books. google. at / books? id = 60 mgmAEACAAJ & printsec = frontcover & hl = de & source = gbs _ ge _ summary _ r & cad = 0 # v = onepage & q & f = false]

Csepreghy-Meznerics, I. (1969) Nouvelles gasteropodes et lamellibranches pour la faune hongroise des gisements tortoniens- inferieurs de la Montagne de Bukk. Annales Historico-Naturales Musei Nationalis Hungarici Pars Mineralogica et Palaeontologica, 61, 63 - 127.

Csepreghy-Meznerics, I. (1972) La faune Tortonienne-inferieure des gisements tufiques de la Montagne de Bukk: Gastropodes II. Az Egri Muzeum Evkonyve (Annales Musei Agriensis), 8, 26 - 46.

Fedosov, A., Puillandre, N., Herrmann, M., Kantor, Yu., Oliverio, M., Dgebuadze, P., Modica, M. V. & Bouchet, P. (2018) The collapse of Mitra: molecular systematics and morphology of the Mitridae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 183, 253 - 337. https: // doi. org / 10.1093 / zoolinnean / zlx 073

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Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 6. A1–A2. Calcimitra bellardii (Hoernes & Auinger, 1880), Coşteiu de Sus (Romania), SFM 12.2243a. B1–B2. Calcimitra bellardii (Hoernes & Auinger, 1880), Coşteiu de Sus (Romania), SFM 12.2243a. C1–C2. Calcimitra bellardii (Hoernes & Auinger, 1880), Coşteiu de Sus (Romania), SFM 12.2243a. D1–D2. Calcimitra bellardii (Hoernes & Auinger, 1880), NHMW 2020/0133/0001, Lăpugiu de Sus (Romania). E1–E2. NHMW 1846/0037/0107a, Calcimitra bellardii (Hoernes & Auinger, 1880), paralectotype, Baden (Austria). F1–F2. NHMW 2020/0135/0001, Calcimitra bellardii (Hoernes & Auinger, 1880), lectotype, Baden (Austria).