Domiporta amoena, Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 27-28

publication ID	https://doi.org/ 10.11646/zootaxa.4983.1.1
publication LSID	lsid:zoobank.org:pub:6A4778D6-195A-4AB1-AA1E-7D8000185B28
DOI	https://doi.org/10.5281/zenodo.5044082
persistent identifier	https://treatment.plazi.org/id/A82A87E9-8A0C-3856-FF4D-F91FFC81F85E
treatment provided by	Plazi
scientific name	Domiporta amoena
status	sp. nov.

Treatment

Domiporta amoena View in CoL nov. sp.

Figs 7A View FIGURE 7 1 –A View FIGURE 1 2 View FIGURE 2 , B 1 –B View FIGURE 1 2 View FIGURE 2 , C 1 –C View FIGURE 1 2 View FIGURE 2 , D 1 –D View FIGURE 1 2 View FIGURE 2 , 8C View FIGURE 8 1 –C View FIGURE 1 2 View FIGURE 2 , D 1 –D View FIGURE 1 2 View FIGURE 2 , E 1 –E View FIGURE 1 2 View FIGURE 2 , F 1 –F View FIGURE 1 2 View FIGURE 2

Mitra fusiformis Brocchi View in CoL — Neugeboren 1860 (pars): 10 [non Brocchi, 1814].

Mitra (Mitra) citima Bell. — Boettger 1906: 7 [non Bellardi, 1887a].

Mitra (Tiara) scrobiculata ( Brocchi, 1814) — Caze et al. 2010: 33, fig. 5C1–C2 [non Brocchi, 1814].

Mitra (Tiara) orientalis var. latisulcata n. var. —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 156 (pars), pl. 41, figs 12–14 [non Kojumdgieva in Kojumdgieva & Strachimirov, 1960].

Type material. Holotype: NHMW 2020 View Materials /0106/0001, SL: 27.7 mm, MD: 8.1 mm, Lăpugiu de Sus ( Romania), figs 7B 1 –B 2, 8D 1 –D 2 . Paratypes: NHMW 2020 View Materials /0106/0002, SL: 29.7 mm , MD: 9.4 mm, Lăpugiu de Sus (Romania), figs 7A 1 –A 2, 8C 1 –C 2 ; NHMW 1868 View Materials /0001/0403, SL: 28.3 mm , MD: 7.9 mm, Lăpugiu de Sus ( Romania), figs 7C 1 –C 2 ; NHMW 2020 View Materials /0106/0003, SL: 34.6 mm , MD: 9.1 mm, Lăpugiu de Sus ( Romania), figs 7D 1 –D 2 ; NHMW 2020 View Materials /0106/0004, SL: 32.5 mm , MD: 9.4 mm, Lăpugiu de Sus ( Romania), figs 8E 1 –E 2 , NHMW 2020 View Materials /0106/0005, SL: 29.8 mm , MD: 8.5 mm, Lăpugiu de Sus ( Romania), figs 8F 1 –F 2 .

Additional material. NHMW 2020 View Materials /0107/0001, 2 specimens , Bad Vöslau ( Austria) ; NHMW 1870 View Materials /0033/0031, 30 specimens , NHMW 1973 View Materials /1615/0087, 10 specimens , NHMW 2016 View Materials /0177/0815, 12 specimens ; NHMW 2016 View Materials /0177/0814, 5 specimens ; NHMW 2016 View Materials /0177/0810, 73 specimens ; NHMW 2016 View Materials /0177/0811, 121 specimens ; NHMW 2020 View Materials /0108/0001, 304 specimens ; NHMW 2016 View Materials /0177/0816, 4 specimens , all 559 Lăpugiu de Sus (Romania) ; NHMW 2020 View Materials /0109/0001, 1 specimen , Nemeşeşti ( Romania) ; NHMW 1867 View Materials /0019/0037, 16 specimens , Coşteiu de Sus ( Romania) .

Type locality. Lăpugiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian.

Etymology. From Latin amoena (= lovely).

Diagnosis. Domiporta species of medium size, slender fusiform profile, sculpture of broad, flat spiral bands or cords separated by deep, rectangular grooves; three spiral cords on spire whorls, split by intercalations of secondary spiral grooves in variable manner, last whorl with broader band along periphery in most specimen, spiral bands on last whorl frequently split by secondary spiral grooves; grooves coincide with intense dark stripes under UV light.

Description. Shell medium sized, slender fusiform with weakly coeloconoid spire and narrowly canaliculate suture. Protoconch high conical of about three smooth, moderately convex whorls. Teleoconch of nine whorls. Early teleoconch whorls subcylindrical, forming high conical spire. Later spire whorls becoming weakly convex and wider. Last whorl high, elongate, with moderately convex periphery placed slightly above adapical termination of aperture, slowly contracting. Sculpture on first three teleoconch whorls cancellate with two blunt spiral cords, each bifurcated by a secondary spiral groove. Axial sculpture and abapical secondary spiral groove fade out on fourth to fifth whorls; spiral cords become broader and flatter, separated by narrower interspaces rectangular in cross-section. Last two spire whorls bearing three broad spiral cords, frequently bifurcated by secondary spiral grooves. Slightly broader spiral cord typically along periphery, often subdivided by secondary spiral grooves. Number of spiral cords on last whorl ranging around 15 depending on number of bifurcations. Spiral interspaces becoming more prominent on base and fasciole, showing delicate axial growth lines. Aperture narrow to moderately narrow, elongate, posteriorly narrowly angulated with very weak parietal swelling. Columellar callus narrow, sharply delimited, bearing three oblique columellar folds; fourth abapical fold strongly reduced. Outer lip thin, without denticles or lirae. Siphonal canal long, wide, weakly twisted, with deeply incised siphonal notch.

Colour pattern in UV light intense, consisting of dark stripes coinciding with spiral grooves and light spiral bands coinciding with spiral cords (figs 7A–C). Dark stripes alternating in width.

Shell measurements and ratios. SL = 20.7–35.0 mm, MD: 6.6–9.4 mm; AA = 28–32°, SL/MD: 3.5–3.8, AL/ AW: 5.3–5.9, AH/S: 2.0–2.4.

Discussion. The species is variable in spiral sculpture ranging from specimens with a distinct broad spiral band along the periphery of the last whorl to specimens with regularly-spaced spiral cords. These morphotypes are connected by numerous intermediate specimens. Domiporta amoena nov. sp. is very abundant at Lăpugiu de Sus ( Romania), documented by hundreds of specimens. These were lumped together with other fusiform Mitridae and labelled as ‘ Mitra scrobiculata ’ in the collection of the NHMW. Boettger (1906) identified this species from Coşteiu de Sus as Mitra citima Bellardi, 1887 . The holotype of Mitra citima , from the middle Miocene of Albugnano ( Italy), however, differs in its more scalate spire, but better preserved material is required to characterise the Italian species (see Ferrero-Mortara et al. 1981, pl. 48, figs 9a–b).

Palaeoenvironment. The clay of the Baden Formation formed in middle to outer neritic settings with up to 250 m water depth ( Hohenegger et al. 2008).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Baden, Bad Vöslau ( Austria); Făget Basin: Nemeşeşti, Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Boettger 1906); Dacian Basin: Opanec ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).