Episcomitra brusinai ( Hoernes & Auinger, 1880 )

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 19-20

publication ID	https://doi.org/10.11646/zootaxa.4983.1.1
publication LSID	lsid:zoobank.org:pub:6A4778D6-195A-4AB1-AA1E-7D8000185B28
DOI	https://doi.org/10.5281/zenodo.5044171
persistent identifier	https://treatment.plazi.org/id/A82A87E9-8A04-385E-FF4D-FA65FE8CFBBD
treatment provided by	Plazi (2021-06-10 00:51:57, last updated 2024-11-24 21:02:59)
scientific name	Episcomitra brusinai ( Hoernes & Auinger, 1880 )
status

Treatment

Episcomitra brusinai ( Hoernes & Auinger, 1880) View in CoL

Figs 5A View FIGURE 5 1 –A View FIGURE 1 2 View FIGURE 2

[ Mitra ] Brusinae nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Brusinae View in CoL nov. form.— Hoernes & Auinger 1880: 76, pl. 9, figs 1a–1b [non fig 2 = Episcomitra leopoldiana View in CoL nov. sp.]. [ Mitra View in CoL ] Brusinae— Cossmann 1899: 156.

Episcomitra fusiformis ( Brocchi, 1814) View in CoL — Landau et al. 2013: 209, pl. 22, fig. 2 [non Brocchi, 1814].

non Mitra brusinae Hoernes & Auinger 1880 View in CoL — Strausz 1966: 363, pl. 41, fig. 19 [=? Episcomitra missile View in CoL nov. sp.].

non Mitra ambigua View in CoL var. Brusinae R. Hoern. View in CoL i Auinger— Friedberg 1911: 13, text-fig. 4 [= unidentifiable fragment].

non M [itraria]. (M [itraria].) brusinae View in CoL (R. Hörn, et Au.)— Sieber 1958a: 153 [= Episcomitra leopoldiana View in CoL nov. sp.].

non Mitraria View in CoL (M.) brusinae (R. Hörn. Auing.) View in CoL — Sieber 1958b: 149 [= Episcomitra leopoldiana View in CoL nov. sp.].

non Mitra brusinae R. Hoern. et Auing. View in CoL — Eremija 1959: pl. 1, figs 3–3a.

Type material. Lectotype (designated herein): NHMW 1854 View Materials /0035/0100, SL: 76.7 mm, MD: 23.2 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 1a–b), figs 5A 1 –A 2.

Revised description. Shell large, moderately slender biconic-fusiform with high spire, elongate last whorl, impressed suture. Protoconch unknown. Teleoconch of eight whorls. Convexity of early teleoconch whorls weak, increasing slightly on later whorls; periphery slightly below mid-whorl. Sculpture of wide-spaced, subobsolete spiral grooves; faint spiral cords on base and fasciole. Last whorl high with periphery above position of adapical termination of aperture, slowly contracting below. Aperture narrow, abapically not constricted. Narrow, thin columellar callus, bearing four wide spaced, oblique columellar folds, weakening abapically. Outer lip thin. Siphonal canal moderately long, wide, straight, with deep siphonal notch.

Shell measurements and ratios. SL = 76.7 mm, MD: 23.2 mm, AA = 28°, SL/ MD: 3.4, AL/AW: 5.9, AH/S: 2.4.

Discussion. Bałuk (1997) was the first who doubted that the specimens described by Hoernes & Auinger (1880) as Mitra brusinai were conspecific and separated fig. 2 as Mitra repleta (= Episcomitra leopoldiana nov. sp.). To solve the status of this species, we select the specimen illustrated by Hoernes & Auinger (1880, pl. 9, fig. 1) as lectotype. Cernohorsky (1976: 378) listed Episcomitra brusinai as a synonym of Isara hoernesi ( Mayer, 1864) , which is clearly incorrect in respect to the completely different shape and much larger size of. E. brusinai .

Hoernes & Auinger (1880) described this species as Mitra Brusinae without explicitly stating after whom it was named. Obviously, the species was dedicated to the Croatian palaeontologist Spiridon Brusina (1845–1908). Therefore, we emend the grammatically incorrect feminine ending to brusinai , as already done by Bałuk (1997: 32).

A specimen from Steinebrunn ( Austria) (NHNW 2020/0123/0001, SL: 74.5 mm, MD: 20.9 mm) differs in its more slender and higher spire and more prominent, twisted fasciole As the intraspecific variability of Episcomitra brusinai is unknown, we refrain from separating the specimen as a distinct species and refer to it in open nomenclature ( Episcomitra cf. brusinai , figs 6B 1 –B 2).

Episcomitra brusinai is similar in shape to several species from the Italian Neogene, but most of those lack spiral sculpture on the last whorl. Episcomitra albigonensis (Bellardi, 1887) from the Pliocene of Italy is almost identical in profile, but is slightly smaller and the spiral sculpture is denser.

Palaeoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1880).

Proto-Mediterranean Sea: Serravallian (middle Miocene): Karman Basin: Pýnarlar Yaylasý, Akpýnar ( Landau et al. 2013).

References

Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]

Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Stamperia Reale, Milano, 2 + 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569

Cernohorsky, W. O. (1976) The Mitridae of the World. Part 1. The Subfamily Mitrinae. Indo-Pacific Mollusca, 3 / 17, 273 - 528. [https: // www. biodiversitylibrary. org / item / 175322 # page / 1 / mode / 1 up]

Cossmann, M. (1899) Essais de Paleoconchologie Comparee. Livraison 3. Cossmann, Lamarre & C., Paris, 201 pp., 8 pls. [https: // www. biodiversitylibrary. org / item / 113486 # page / 421 / mode / 1 up]

Eremija, M. (1959) Palaontologische Neuigkeiten aus Neogenschichten sudlich vor der Stadt Glina (Croatien). Anneles Geologique de la Peninsule Balkanique, 26, 185 - 193.

Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 & 5, 1 - 112, pls. 1 - 5 (1911) & 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]

Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405

Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576

Mayer, K. (1864) Die Tertiar-Fauna der Azoren und Madeiren. Systematisches Verzeichniss der fossilen Reste von Madeira, Porto Santo und Santa Maria nebst Beschreibung der neuen Arten. published by the author, Zurich, vi + 107 pp. [https: // books. google. at / books? id = wS 4 - AAAAcAAJ & pg = PP 11 & source = kp _ read _ button & redir _ esc = y # v = onepage & q & f = false]

Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Sieber, R. (1958 b) Die Tortonfauna von Steinabrunn bei Drasenhofen (Bez. Mistelbach, N. - O.). Verhandlungen der Geologischen Bundesanstalt, 1958, 142 - 155. [https: // opac. geologie. ac. at / ais 312 / dokumente / VH 1958 _ 142 _ A. pdf]

Strausz, L. (1966) Die Miozan-Mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 5. A1–A2. Episcomitra brusinai (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0100. B1–B2. Episcomitra cf. brusinai (Hoernes & Auinger, 1880), Steinebrunn (Austria), NHNW 2020/0123/0001. C1–C2. Episcomitra missile nov. sp., holotype, Guntersdorf (Austria), NHMW 202/0127/0001. D1–D2. Episcomitra missile nov. sp., paratype, Grund (Austria), NHMW 202/0128/0001. E1–E2. Episcomitra friedbergi (Cossmann, 1912), lectotype, Lăpugiu de Sus (Romania), NHMW 20200124/0001. F1–F2. Episcomitra friedbergi (Cossmann, 1912), Lăpugiu de Sus (Romania), NHMW 1868/0001/0398. G1–G2. Episcomitra neubaueri nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0084. H1– H2. Episcomitra neubaueri nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0086. I1–I2. Episcomitra neubaueri nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 1855/0043/0006.