Penstemon linarioides var. coloradoensis (A.Nelson) C.C.Freeman, 2017

Freeman, Craig C., 2017, Nomenclatural novelties and notes in Penstemon (Plantaginaceae), PhytoKeys 80, pp. 33-39 : 34-35

Penstemon linarioides var. coloradoensis (A.Nelson) C.C.Freeman stat. nov.

Basionym.

Penstemon coloradoensis A. Nelson, Bull. Torrey Bot. Club 26: 355. 1899; Penstemon linarioides subsp. coloradoensis (A. Nelson) D.D. Keck.

Type.

USA. COLORADO: Mancos, common on sage plains and foothills throughout the La Platas, 23 Jun 1898, C.F. Baker, F.S. Earle, and S.M. Tracy 70 (lectotype: RM, RM0004187 [JSTOR image]; isolectotypes: CAS, CAS0006620 [CAS website image], F, F0072589F [JSTOR image], GH, GH00091215 [GH website image], K, K000979702 [K website image], KSC, MICH, MICH1108038 [MICH website image], MO, MO-100230852, NDG, NDG45904 [JSTOR image], NEB, NEB-v-0000619 [JSTOR image], NY, NY00091049 [NY website image], OS, OS0000264 [JSTOR image], RM, RM0004188 [JSTOR image], RSA, RSA 0006119 [JSTOR image], RSA 0006120 [JSTOR image], US, US 00122246 [ US website image]; probable isolectotype: NDG, NDG45695 [JSTOR image]) .

In describing Penstemon coloradoensis , Nelson (1899) stated " I have before me specimens from two collections made near Mancos, Colo., by Messrs. Baker, Earle, and Tracy, 1898 and distributed as P. caespitosus Nutt. Also from two collections by Professor Crandall, from Hotchkiss, Colo., 1892, and from Durango, 1898, both distributed as P. linarioides Sileri Gray. " Nelson (1899) did not designate a holotype; the collections he cited are syntypes. Keck (1937) designated C.F. Baker, F.S. Earle, and S.M. Tracy 70 (RM) as the lectotype, of which there currently are two sheets, suggesting a second-step lectotypification might be necessary. Accession labels on the two sheets indicate they were received from the herbaria of F.S. Earle in 1899 (RM0004187; accession number 13703) and George E. Osterhout in 1938 (RM0004188; accession number 165282), subsequent to Keck’s lectotype designation. RM0004187 bears annotations by A. Nelson ( Penstemon coloradoensis A. N. n. sp., without date), F.W. Pennell ( P. coloradoensis , type, 1916), and D.D. Keck ( P. linarioides A. Gray subsp. coloradoensis (A. Nelson) D.D. Keck, type, 1936). RM0004188, the later accession, was not annotated by Nelson, Pennell, or Keck. Whether Nelson had another sheet at RM that was lost, destroyed, or sent elsewhere, or had access to a sheet from some other herbarium, is not known. Of the two sheets currently at RM, only RM0004187 appears to have been there when Nelson described P. coloradoensis and later seen by Keck when he lectotypified the name. RM0004187 is taken as the lectotype, RM004188 is taken as an isolectotype, and no second-step lectotypification is needed. As an aside, the sheet at KSC is P. crandallii A. Nelson var. crandallii .