Bathyphantes paracymbialis, Tanasevitch, Andrei V., 2014

Tanasevitch, Andrei V., 2014, New species and records of linyphiid spiders from Laos (Araneae, Linyphiidae), Zootaxa 3841 (1), pp. 67-89 : 73-75

Bathyphantes paracymbialis new species

Figs 15–23

Types. Holotype. ♂ LAOS, Luang Prabang Prov., SE Luang Prabang, Nam Khan, Xieng Ngeun Distr., Ban Keng Koung , 372 m a.s.l., 19°40'963''N 102°18'442''E, disturbed forest, valley, waterfall, dry stream bed, sieving and Winkler, 21.–23.II.2008, leg. P. Jäger . Paratypes. 1♂, 1♀, Luang Nam Tha Prov., Vieng Phou Kha, Phou Pasat, Tham Pasat Thia, 705 m a.s.l., 20°46'37.2''N 101°01'00.2''E, outside cave, secondary forest, leaf litter, sieving, 4.III.2008, leg. P. Jäger . 5♀ ( MHNG), Oudomxai Prov., ca 3 km E of Tad Lak 11, SE of Oudomxai (= Muang Xai) City, 20º36'20''N 102º01'16''E, 1020 m a.s.l., 18.IX.2010, leg. P. Schwendinger [ LT-10/12 ] . 1♂, 3♀ ( MHNG), THAILAND, Chiang Mai Prov. and Distr., Doi Suthep , 960 m a.s.l., pitfall traps, 28.IV.–30.V.1987, leg. P. Schwendinger . 1♂, 1♀ ( MHNG), same site, 30.V.–4.VII.1987, leg. P. Schwendinger . 1♀ ( MHNG), same site, 30.VII.–30.VIII.1986, leg. P. Schwendinger . 1♀ ( MHNG), Doi Suthep , 1180 m a.s.l., pitfall traps, 2.VII.–2.VIII.1987, leg. P. Schwendinger . 1♀ ( MHNG), same site, 28.IV.–30.V.1987, leg. P. Schwendinger . 1♂, 1♀ ( MHNG), Doi Suthep , 1155 m a.s.l., pitfall traps, 10.I.–11.II.1997, leg. P. Schwendinger . 1♂ ( MHNG), same site, 14.XII.–10.I.1997, leg. P. Schwendinger . 1♀ ( MHNG), Doi Suthep , 1150 m a.s.l., 30.XI.1996, leg. P. Schwendinger . 1♂, 1♀ ( MHNG), same site, 1150 m a.s.l., pitfall traps, 14.XII.1996.– 10.I.1997, leg. P. Schwendinger . 1♀ ( MHNG), Doi Chiang Dao , 510 m a.s.l., pitfall traps, 22.XII.1990.– 15.I.1991, leg. P. Schwendinger . 1♂ ( MHNG), Chomthong Distr., Doi Inthanon , Pha Mon Valley , 1050 m a.s.l., 23.XI.1996, leg. P. Schwendinger . 1♂, 2♀ ( MHNG), Fang Distr., Doi Ankhang , 1500 m a.s.l., 30.X.1987, leg. P. Schwendinger . 1♂ ( MHNG), Chiang Rai Prov., Wiang Pa Pao Distr., Ban Huay Ya Sai , 780 m a.s.l., 28.I.1988, leg. P. Schwendinger. 1♀ ( MHNG), Phetchabun Prov., Nam Nao N.P. , 850 m a.s.l., 14.IX.1990, leg. P. Schwendinger . 1♀ ( MHNG), Ranong Prov., Kapoe Distr ., Khlong Nakha Wildlife Sanctuary , near Headquarters, 30 m a.s.l., 29.I.1991, leg. P. Schwendinger . 1♀ ( MHNG), Pattani Prov., Nam Tok Sai Khao N.P. , ca. 20 km NW of Yala City, Sai Khao Waterfall , 260–350 m a.s.l., evergreen rain forest near a stream, 23.X.1999, leg. P. Schwendinger [ THA-99/5 ]. 1♀ ( MHNG), WEST MALAYSIA, Pahang, Genting Highlands , 1650 m a.s.l., primary forest, steep slope, sieving, 5.IV.1993, leg. I. Löbl & F. Calame [ WM93-29 ].

Etymology. The species name refers to the peculiar shape of the paracymbium; adjective.

Diagnosis. The new species is very similar to B. floralis Tu & Li, 2006 and to the Japanese B. robustus Oi, 1960 , but the male can easily be distinguished by the presence of a swollen outgrowth at middle part of the paracymbium and the smaller embolic coil. The female differs by a short but clearly developed scapoid, while in B. floralis and B. robustus the scapoid looks like a small knob, as well as by the loops of the entrance ducts in B. paracymbialis which are wider than in the other two species.

Description. Male (paratype from Tham Pasat Thia, Laos). Total length 1.93. Carapace unmodified, 0.88 long, 0.70 wide, greyish pale brown. Eyes normal. Chelicerae 0.35 long. Legs pale brown, yellow. Leg I, 3.87 long (0.98+0.23+1.03+0.98+0.65), IV, 3.89 long (1.05+0.20+1.03+0.98+0.63). Chaetotaxy. TiI: 2-1-1-0, II: 2-0-1-0, III-IV: 2-0-0-0; metatarsi spineless. Metatarsi I–III with a trichobothrium each. TmI, 0.26. Palp ( Figs 15–19 ): Paracymbium with a swollen outgrowth at middle part. Lower part of tegulum with a thick process directed backward. Suprategulum with two apophyses: one as a narrow stripe bifurcated apically directed forward, other one wide and rounded directed downward. Embolic plate (sensu Beatty & al. 1991; Saaristo 1995) (= lamella or lamella characteristica auct.) large, flat, with dorsad projection proximally. Embolus relatively thick and short, coiled. Opisthosoma 1.03 long, 0.48 wide, dorsally with a buckling in first third. Colouration: anterior part of opisthosoma pale, posterior part grey with indistinct pale transverse stripes.

Female (paratype from Oudomxai City). Total length 2.00. Carapace unmodified, 0.90 long, 0.65 wide, greyish pale brown. Eyes normal. Chelicerae 0.40 long. Legs pale brown, yellow. Leg I, 3.91 long (1.00+0.25+1.05+0.98+0.63), IV, 3.79 long (1.00+0.25+1.03+0.88+0.63). Chaetotaxy. FeI: 1-1-0-0, II: 1-0-0-0, III-IV: 0-0-0-0; TiI: 2-1-1-0, II: 2-0-1-0, III-IV: 2-0-0-0; metatarsi spineless. Metatarsi I–III with a trichobothrium each. TmI, 0.23. Opisthosoma 1.08 long, 0.68 wide, dorsally without buckling, grey to dark grey, in some specimens with indistinct pattern of pale transverse stripes. Epigynum ( Figs 20–23 ): Anterior wall with a small conical scapoid (= scapus, auct.), posterior median plate with a narrow parmula. Entrance ducts making two wide loops. Receptacula almost spherical.

Variation. Body and leg coloration vary from pale to dark; the paracymbial outgrowth can be pointed to rounded, edge of scapoid can be transparent, which makes it shape visually different from the depicted in Figs 20–22 .

Distribution. Laos, Thailand, West Malaysia.

Figures

FIGURES 15 – 23. Bathyphantes paracymbialis n. sp. male paratype (from Tham Pasat Thia, Laos, 15 – 19) and female paratypes (from Oudomxai City, 20 – 23). 15, Left palp, retrolateral, 16, prolateral. 17, 18, Paracymbium, lateral, different aspects. 19, Distal suprategular apophysis, lateral. 20 – 22, Epigynum scapoid variety, ventral. 23, Epigynum, ventral, cleared.