Thia scutellata (Fabricius, 1793), 1962

BOYKO, CHRISTOPHER B., 2002, A Worldwide Revision Of The Recent And Fossil Sand Crabs Of The Albuneidae Stimpson And Blepharipodidae, New Family (Crustacea: Decapoda: Anomura: Hippoidea), Bulletin of the American Museum of Natural History 2002 (272), pp. 1-396 : 396

Thia scutellata (Fabricius, 1793) Figure 1A

Hippa scutellata Fabricius, 1793: 474–475 . – Weber, 1795: vii. – Holthuis, 1962: 125–128, pl. 4. – Zimsen, 1964: 649.

Albunea scutellata: Fabricius, 1798: 397 . – Herbst, 1804: 31 (list). – Desmarest, 1823: 284. – Desmarest, 1825: 173–174.

albunea scutellata: Latreille, 1803: 172 . – Lamarck, 1818: 224.

Lepidopa scutellata: Ortmann, 1896: 226 (part). – Gordon, 1938: 188 (part).

Lepidopa scutellata: Holthuis, 1962: 125–128 .

Thia scutellata: Holthuis, 1962: 125–128 . – Christiansen, 1969: 40–41, fig. 14.

Thia scutella [sic]: Salva and Feldmann, 2001: 24, fig. 12a–c.

not Albunea scutellata: H. Milne Edwards, 1837b: 204 , pl. 21, figs. 9–13. – Chenu and Desmarest, 1877: 32, fig. 22 (= Lepidopa benedicti Schmitt, 1935 ).

not Albunea scutellata: Gibbes, 1850b: 187 (=

Lepidopa websteri Benedict, 1903 ). not Albunaea [sic] scutellata: Dana, 1852: 406 (=

Lepidopa chilensis Lenz, 1902 ). not Lepidopa scutellata: Stimpson, 1858: 230 .—

Stimpson, 1859: 79. – Faxon, 1895: 237 (list).

– Ortmann, 1896: 226 (part), 227 (part). – Gor­

don, 1938: 188 (list, part) (= Lepidopa rich­

mondi Benedict, 1903). not Lepidops [sic] scutellata: Miers, 1878: 332

(part) (= Lepidopa chilensis Lenz, 1902 ). not Lepidops [sic] scutellata: Miers, 1878: 332

(part). – Moreira, 1901: 30, 88–89 (= Lepidopa

richmondi Benedict, 1903). not Lepidopa scutellata: Ortmann, 1896: 226

(part). – Benedict, 1903: 894, fig. 6*. – Schmitt,

1935: 209–210* (= Lepidopa benedicti

Schmitt, 1935). not Lepidopa scutellata: Ortmann, 1896: 227

(part) (= Lepidopa websteri Benedict, 1903 ). not Lepidopa scutellata: Ortmann, 1896: 227

(part). – Gordon, 1938: 188 (part) (= Lepidopa

chilensis Lenz, 1902). not Lepidopa scutellata: Schmitt, 1924: 96 (=

Lepidopa sp. indet.).

REMARKS: This species was the last albuneid to be recognized as a brachyuran and removed from the Albuneidae (Holthuis, 1962) . The longstanding confusion about the identity of this species dates back to Weber (1795), who placed it under the heading ‘‘Cancroidea dubia.’’ This species lat­ er was designated the type of the genus Lepidopa by Stimpson (1858), who incorrectly applied the name to a species of that genus. The type species of Lepidopa was subsequently designated as L. venusta by the plenary powers of the ICZN (ICZN, 1964). The type(s) of Hippa scutellata are lost (Holthuis, 1962; Zimsen, 1964) and a neotype (RMNH D351) was designated by Holthuis (1962). The familiar placement of the species has been recently verified by Salva and Feldmann (2001). This specific name is no. 1956 on the ‘‘Official list of specific names in zoology’’ (ICZN, 1964).

This species is also adapted to a sand­dwelling mode of life and has been collected in the same samples as Albunea carabus in shallow (3–7 m) water off Spain (Rubió and Holthuis, 1976).

Figures

Fig. 1. Convergent crabs: A. Thia scutellata (Fabricius, 1793). B. Corystes cassivelaunus (Pennant, 1777). C. Albunea carabus (Linnaeus, 1758). D. Ranina ranina (Linnaeus, 1758). E. Notopus dorsipes (Fabricius, 1787). A, B, after Hayward et al. (1995), C, after Zariquiey Alvarez (1968), D, after Tinker (1965), E, specimen from Madagascar (MNHN). Figures not given to scale.