Amara (Curtonotus) harpaloides Dejean, 1828

Li, Yihang, Li, Haoyuan & Shi, Hongliang, 2024, Revision of the macropterous subgenus Curtonotus from east China, with the description of a new species (Carabidae, Zabrini, Amara), ZooKeys 1190, pp. 39-73 : 39

Amara (Curtonotus) harpaloides Dejean, 1828

Fig. 10

Amara harpaloides Dejean, 1828: 514 (type locality: “Sibirien” [= Barnaul, Altai Krai, Russia]; syntypes in MNHN); Hieke 1999: 162.

Curtonotus convexicollis Putzeys, 1866: 232 (type locality: “Siberia”; holotype in MNHN); synonymized by Hieke 1999: 162.

Specimens examined.

2 males (CLYH), China, Hebei, Zhangjiakou, Zhuolu, Lingshan Scenic Spots , 40.054300°N, 115.487502°E, 1788 m, 2021.08.02, Yihang Li leg. ; 1 female (CLYH), China, Beijing, Songshan National Reserve , 40.534820°N, 115.7541325°E, 1380 m, 2022.08.07, Yihang Li leg. ; 1 male (CLHY), China, Beijing, Qingshui Town, Hongkou Village , light trap, 39.99464407°N, 115.48366919°E, 950 m, 2022.07.24, Haoyuan Li leg. ; 19 males, 17 females (CBJFU), China, Inner Mongolia, Genhe, Greater Khingan Ecological Station , 50.8061°N, 121.5824°E, 726 m, 2018.08.28, Hongliang Shi leg. ; 1 male (CLYH), China, Hebei, Zhangjiakou, Hailiutu, Dayuedai , 41.176428°N, 114.512037°E, 1390 m, 2022.09.11, Cong Wang leg. ; 1 female (CBJFU), China, Gansu, Yongdeng, Liancheng Township , 36.5940°N, 102.8326°E, 1927 m, 2021. 08.04, Youyan Huang & Hanshuo Liu leg. ; 1 female (CBJFU), China, Qinghai, Haixi, Tianjun County, Kuaier’ma, 37.4724°N, 98.7772°E, 3765 m, 2022.08.05, Hongliang Shi leg. ; 3 males, 11 females (CBJFU), China, Qinghai, Zekong county, Maixiu forestry center, 35.2706°N, 101.9304°E, 2962 m, 2019.08.23, Weifeng Yan leg.

Chinese common name.

婪暗步甲.

Diagnosis.

Medium to large-sized species, BL = 10.0-12.0 mm; body black, legs dark brown to black; head small, ~ 1/2 of pronotum maximum width, with two supraorbital setae. Pronotum (Fig. 10B ) subcordate, widest near middle, densely punctate at basal region, sparsely punctate at mid-anterior region; lateral margins weakly sinuate or nearly straight on posterior half; posterior angles more or less protruded laterally, nearly rectangular, apex without denticulate. Elytra oblong, widest near middle; finely punctate on basal half of striae; humeral tooth strongly and straightly protruded; elytra with isodiametric microsculpture, stronger in females; lateral sides of abdominal sternites wrinkled. Male mesotibiae projection composed of two denticles (Fig. 10C ); proximal denticle acutely pointed, near apical third of tibiae; distal denticle slightly smaller and wider than proximal one, between the proximal one and tibiae apex. Male genitalia with slightly long apical lamella (Fig. 10F ), straight and widely triangular, gradually narrowed to apex, apex rounded; gonocoxite 2 (Fig. 10G ) of ovipositor elongate, length ~ 1.5 × greatest width, apex widely rounded.

Comparison.

Among this species group, A. (C.) harpaloides is most similar to A. (C.) macronota and A. (C.) beijingensis , and can be distinguished from these two by having smaller elytra punctures, stronger elytral microsculpture in females, shallowly sinuate pronotum lateral margins before the posterior angles, and humeral tooth more strongly protruded.

Distribution.

China (Gansu, Hebei, Beijing, Heilongjiang, Qinghai, Sichuan, Shanxi, Inner Mongolia, Sichuan), Russia (west Siberia, east Siberia, Far East).

Figures

Figure 10. Amara (Curtonotus) harpaloides A dorsal habitus, male (Zhangbei, Hebei) B pronotum posterior angle (Tianjun, Qinghai) C male mesotibia D lateral view of aedeagus E right paramere F apical lamella G female gonocoxite (Yongdeng, Gansu). Scale bar: 5 mm (A); 1 mm (B-E); 0.5 mm (F, G).