Tomarus roigjunenti Neita & Ratcliffe, 2017

López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 69-70

Tomarus roigjunenti Neita & Ratcliffe, 2017

( Figs. 9K , 16O , 20F , 28F ; 47 )

Tomarus roigjunenti Neita & Ratcliffe, 2017: 8 . Original combination.

Male holotype not examined. Male paratype ( USNM) “ Neuquen / Capital Federal / 1.iii.1961 / Dafuern // TOMARUS / ROIGJUNENTI / NEITA & RATCLIFFE / PARATYPE ”. Female paratype ( USNM) “ Neuquen / Capital / 1.iii.1961 // TOMARUS / ROIGJUNENTI / NEITA & RATCLIFFE / PARATYPE ”. Two female paratypes ( USNM) “ Mendoza / C.S. Reed // TOMARUS / ROIGJUNENTI / NEITA & RATCLIFFE / PARATYPE ” Type locality: Neuquén, Argentina .

Description. Habitus as in Fig. 28F . Length 26.3–27.6 mm; humeral width 12.6–13.9 mm. Color dark reddish brown to black. Head: Frons coarsely rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus narrowed towards apex, base 3.8 wider than apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Interocular distance 4 times an eye width. Pronotum: Surface with dense, large punctures. Apical tubercle small, rounded. Fovea shallow, striate, narrow (1/5 as wide as interocular distance). Scutellum: Surface with minute, sparse punctures. Elytra: First interval punctate, punctures similar in size to other intervals; sutural stria complete. Inner surface of apex with rounded, small tubercles forming 38–40 parallel lines. Abdomen: Apex of tergite IV with a triangular wide area with large, elongate tubercles forming about 18 diagonal, parallel lines. Pygidium with dense rugosity on basal 3rd ( Fig. 9K ), apex regularly rounded. Venter: Apex of prosternal process flat, longitudinally oval. Metasternum with short, sparse setae on anterior angles. Legs: Protibia tridentate with an additional inconspicuous, basal convexity. Metatibia triangular; apex slightly crenulate, with 21–25 spinules. Male genitalia: Spiculum gastrale without basal plates. Phallobase 1.1 times longer than parameres. Parameres with lateral teeth large, acute; apical half narrowed, apices outwards, rounded ( Figs. 16O , 20F ). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 1–3 spine-like, accessory lamellae, with dense granules at base.

Diagnosis. Tomarus roigjunenti can be recognized by the following character combination: frontal tubercles transverse; pronotal fovea and tubercle inconspicuous ( Fig. 28F ); pronotum deeply and densely punctate; metasternum with sparse, minute setae on anterior angles; internal sac with a complex of 1–3 spine-like, accessory lamellae; and parameres with lateral teeth large and long, arcuate; apical half narrowed, apices outwards, rounded ( Figs. 16O , 20F ).

Distribution. Argentina ( Neita-Moreno & Ratcliffe 2017).

Locality records ( Fig. 47 ). 4 examined specimens from USNM. Records from Neita-Moreno & Ratcliffe (2017). ARGENTINA (20). Chubut (1): Chubut. Buenos Aires (1). Buenos Aires. Mendoza (4). Tunuyan; Uspallata. Neuquén (9). Capital Federal ; Barrancas ; Collón Cura ; Covunco ; Isla Victoria ; Laguna Blanca ; Vatra Lauquen. Río Negro (5). Allen ; Cipolletti ; General Roca ; Río Negro ; Villa Regina .

Natural history. Some specimens were found in sand dunes and adults are attracted to lights at night (NeitaMoreno & Ratcliffe 2017).

Figures

FIGURE 9. Pygidium of male. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) cuniculus, D) L. (Ligyrus) fossor, E) L. (Ligyrus) paranaensis, F) Tomarus bituberculatus, G) T. colombianus, H) T. maracaiboensis, I) T. pilcopataensis, J) T. pullus, K) T. roigjunenti, L) T. rostratus.

FIGURE 16. Parameres in frontal view. A) Tomarus adoceteus, B) T. amazonicus, C) T. bituberculatus, D) T. colombianus, E) T. discrepans, F) T. gyas, G) T. laevicollis, H) T. laticaudus, I) T. maimon, J) T. maracaiboensis, K) T. maternus, L) T. pilcopataensis, M) T. pullus, N) T. pumilus, O) T. roigjunenti, P) T. rostratus, Q) T. selanderi, R) T. subtropicus.

FIGURE 20. Parameres in lateral view. A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus.

FIGURE 28. Habitus of the species of Tomarus: A) Tomarus maracaiboensis, B) T. maternus, C) T. pilcopataensis, D) T. pullus, E) T. pumilus, F) T. roigjunenti, G) T. rostratus, H) T. selanderi, I) T. subtropicus. Scale bar: 5 mm.

FIGURE 47. Distributional records of Tomarus pilcopataensis, T. roigjunenti, and T. pumilus.