Tomarus selanderi ( Cartwright, 1959 )

López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 70-71

publication ID

https://doi.org/ 10.11646/zootaxa.5211.1.1

publication LSID

lsid:zoobank.org:pub:5754769C-B747-4714-BDD9-7D5509D48BEB

DOI

https://doi.org/10.5281/zenodo.7345129

persistent identifier

https://treatment.plazi.org/id/A209493B-FD1A-FFC7-AFA6-1F8DFE16B9EA

treatment provided by

Plazi

scientific name

Tomarus selanderi ( Cartwright, 1959 )
status

 

Tomarus selanderi ( Cartwright, 1959)

( Figs. 2J View FIGURE 2 , 4H View FIGURE 4 , 6F View FIGURE 6 , 14G View FIGURE 14 , 16Q View FIGURE 16 , 20H View FIGURE 20 , 28H View FIGURE 28 ; 45 View FIGURE 45 )

Bothynus selanderi Cartwright, 1959: 537 . Original combination.

Male holotype (USNM) “ MEXICO / Los Mochis / Sinaloa // at light / 22 July 1955 // RB&JM Selander // Type No / 63912 / USNM // HOLOTYPE / Bothynus / selanderi / Cartwright”. Female allotype (USNM) “ MEXICO / Los Mochis / Sinaloa // at light / 22 July 1955 // RB&JM Selander // Type No / 63912 / USNM // ALLOTYPE / Bothynus / selanderi / Cartwright”. Twelve male and 21 female paratypes (USNM) “ MEXICO / Los Mochis / Sinaloa // at light / 22 July 1955 // RB&JM Selander // Type No / 63912 / USNM // PARATYPE / Bothynus / selanderi / Cartwright”. “ MEXICO / Mazatlán / Sinaloa // 21 July 1955 // RB&JM Selander // Type No / 63912 / USNM // PARATYPE / Bothynus / selanderi / Cartwright”. Male paratype (MNHN) “Los Mochis / Sinaloa, Mex / 22-VII-1955 // R&J Selander // PARATYPE / Bothynus / selanderi / Cartwright // PARATYPE // MNHN / EC6732 About MNHN ”. Male paratype (MNHN) “Los Mochis / Sinaloa, Mex / 22-VII-1955 // R&J Selander // PARATYPE / Bothynus / selanderi / Cartwright // PARATYPE // MNHN / EC6733 About MNHN ”. Type locality: Los Mochis, Sinaloa, Mexico.

Description. Habitus as in Fig. 28H View FIGURE 28 . Length 21.0–30.0 mm; humeral width 10.1–14.5 mm. Color dark reddish brown to black. Head: Frons and clypeus coarsely and densely rugose. Frontoclypeal region with 2 conical tubercles separated by about 4.5–5.0 tubercle diameters. Clypeus narrowed towards apex, base 2.5 times wider than apex. Clypeal teeth triangular, separated by a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed ( Fig. 4H View FIGURE 4 ). Interocular distance equal to 3.5 times an eye width. Pronotum: Surface with small punctures, punctures denser and deeper on anterior and posterior angles. Apical tubercle prominent, acute, slightly recurved. Subapical fovea deep, rounded, as wide as interocular distance or slightly narrower, equal in both sexes, surface transversely rugose. Scutellum: Surface with minute, sparse punctures. Elytra: Punctures on first interval as large as those on other intervals. Inner surface of apex with rounded, large tubercles forming 17 parallel lines. Abdomen: Apex of tergite IV with a triangular wide area with large, rounded tubercles forming 11–12 diagonal, parallel lines. Pygidium with dense rugosity on basal 3rd. Apex rounded. Venter: Apex of prosternal process flat, longitudinally oval to subrectangular. Metasternum with minute setae. Legs: Protibia tridentate, sometimes with a basal small denticle. Apex of metatibia crenulate, with 13–15 spinules. Male genitalia: Spiculum gastrale without basal plates ( Fig. 14G View FIGURE 14 ). Phallobase 1.1 times longer than parameres. Parameres without dorsal teeth, apex wide, triangularly dilated ( Figs. 16Q View FIGURE 16 , 20H View FIGURE 20 ). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 5 spine-like, accessory lamellae, with dense granules at base.

Diagnosis. Tomarus selanderi can be differentiated by the following character combination: frontoclypeal tubercles conical; pronotal tubercle acute and prominent ( Fig. 28H View FIGURE 28 ); pronotal fovea shallow, as wide as or narrower than interocular distance; protibia tridentate, rarely with an additional basal denticle; apex of metatibia with 14–15 spinules; parameres without dorsal teeth; apex wide, triangularly dilated ( Figs. 16Q View FIGURE 16 , 20H View FIGURE 20 ).

Distribution. Tomarus selanderi is an endemic species found along the Pacific coast of Mexico ( Cartwright 1959; Endrödi 1985; Ratcliffe et al. 2013).

Locality records ( Fig. 45 View FIGURE 45 ). 205 examined specimens from CAFESI, EMAC, FSCA, CNIN, IEXA, MNHN, UNSM, USNM. Some data from Ratcliffe et al. (2013). MEXICO (214). Colima (2): Manzanillo. Guerrero (2): Acapulco. Jalisco (17): Careyes; Chamela; Guadalajara; Melaque; Puerto Vallarta. Michoacán (1): Jungapeo. Nayarit (85): Acaponeta; El Venado; Ruiz; San Blas ; 15 km S San Blas; Sayulita; Singayta; Tecuala; Yago. Oaxaca (2): Pinotepa Nacional; Puerto Escondido. Sinaloa (98): El Mármol; Escuinapa; Los Mochis; Mazatlan; Novalato; Piedra Blanca. Sonora (6): Esperanza, Guaymas, Hermosillo .

Natural history. Specimens have been collected between 30–1500 m ( Ratcliffe et al. 2013) in dry forests of Pacific coast of Mexico.

FSCA

Florida State Collection of Arthropods, The Museum of Entomology

CNIN

Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico

MNHN

Museum National d'Histoire Naturelle

UNSM

University of Nebraska State Museum

USNM

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Dynastidae

Genus

Tomarus

Loc

Tomarus selanderi ( Cartwright, 1959 )

López-García, Margarita M. & Deloya, Cuauhtémoc 2022
2022
Loc

Bothynus selanderi

Cartwright, O. L. 1959: 537
1959
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