Tomarus rostratus Dupuis, 2014

López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 70

publication ID

https://doi.org/ 10.11646/zootaxa.5211.1.1

publication LSID

lsid:zoobank.org:pub:5754769C-B747-4714-BDD9-7D5509D48BEB

DOI

https://doi.org/10.5281/zenodo.7345121

persistent identifier

https://treatment.plazi.org/id/A209493B-FD1A-FFC4-AFA6-1871FA3EBAE8

treatment provided by

Plazi

scientific name

Tomarus rostratus Dupuis, 2014
status

 

Tomarus rostratus Dupuis, 2014

( Figs. 5F View FIGURE 5 , 7H View FIGURE 7 , 9L View FIGURE 9 , 13J View FIGURE 13 , 14F View FIGURE 14 , 16P View FIGURE 16 , 20G View FIGURE 20 , 22B View FIGURE 22 , 28G View FIGURE 28 ; 49 View FIGURE 49 )

Tomarus rostratus Dupuis, 2014: 1 . Original combination.

Male holotype (MNHN) “ Peru. / Tomarus / Er. / Maimon // MUSÉUM PARIS / 1952 / coll. R. OBERTHÜR // Tomarus rostratus n. sp. / HOLOTYPE / F. Dupuis 2014 // HOLOTYPE // MNHN / EC4187”. Type locality: Peru.

Description. Habitus as in Fig. 28G View FIGURE 28 . Length 17.4–27.5 mm; humeral width 8.3–10.0 mm. Color dark reddish brown. Head: Frons coarsely and irregularly rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus trapezoidal, not strongly narrowed toward apex; apex with 2 triangular teeth separated by about 1.5 tooth diameters. Mandible with 2 apical teeth and a lateral, widely rounded tooth ( Fig. 5F View FIGURE 5 ). Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Pronotum: Surface punctate, punctures rounded, smaller than punctures on elytra; densely distributed on apical and lateral margins. Apical margin with a small tubercle, visible in lateral view. Pronotum without rugose fovea, with small to large punctures on each side of tubercle. Scutellum: Surface without punctures. Elytra: Sutural stria complete. Elytral punctures dense and deep, punctures mainly ocellate, 3 pairs of distinct double rows. Abdomen: Pygidium convex (male) to flat (female) in lateral view; surface deeply rugopunctate on basal 3rd with disc smooth (male) ( Fig. 9L View FIGURE 9 ), or completely rugopunctate (female). Apical margin of sternite VII straight in male, slightly emarginate with notch broad and shallow in female. Apex of sternite VIII emarginate, notch wide in male and narrow and deep in female. Venter: Prosternal process long, thick, apex transversely oval to rounded, and bordered with long, pale yellow setae. Metasternum with short, sparse setae on anterior angles. Legs: Protibia tridentate with an additional small, basal denticle. Apex of metatibia slightly crenulate, with 8–15 spinules. Female genitalia: Coxite surface strongly concave ( Fig. 13J View FIGURE 13 ). Male genitalia: Spiculum gastrale with basal plates ( Fig. 14F View FIGURE 14 ). Parameres widened at apical 4th, elongate from basal 4th to apex; sides parallel, without teeth or lateral projections ( Fig. 16P View FIGURE 16 ), regularly curved at lateral view ( Fig. 20G View FIGURE 20 ). Internal sac with small copulatory lamella, lamellar spiny belt reduced, complex of accessory lamellae with 6 spines joined at the base by dense granules ( Fig. 22B View FIGURE 22 ).

Diagnosis. Tomarus rostratus is unique in the genus because of its extremely elongate parameres, without lateral teeth. It can be also differentiated by the following combination of characters: clypeus trapezoidal; pronotum with small tubercle and without rugose fovea, with small to large punctures on each side of tubercle ( Fig. 28G View FIGURE 28 ); protibia tridentate with a small denticle; apical border of pygidium strongly truncate in male; apical margin of sternite VIII emarginated in female; coxite surface strongly concave ( Fig. 13J View FIGURE 13 ); spiculum gastrale with basal plates ( Fig. 14F View FIGURE 14 ); internal sac of aedeagus with small copulatory lamella and reduced lamellar spiny belt ( Fig. 22B View FIGURE 22 ).

Distribution. Tomarus rostratus is an endemic species from the coastal desert of Peru ( Dupuis 2014) and Chile ( López-García & Deloya 2018).

Locality records ( Fig. 49 View FIGURE 49 ). 39 examined specimens from FSCA, MEKRB, MNHN, USNM, SENASA, UGBC. CHILE (12). Arica y Parinacota: Cuya; Puente Chapa (50 km S Arica); Valle de Azapa (6 km E Arica). PERU (27). Arequipa (3): Camaná, Quilca; Quiroz. Lima (19): Canta, Santa Rosa de Quives; Cañete; Huarochirí, Santa Eulalia; Huaura, Végueta, Medio Mundo; Lima. No data (5): Peru.

Natural history. Specimens have been found between 12–1,036 m elevation between August and February.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Dynastidae

Genus

Tomarus

Loc

Tomarus rostratus Dupuis, 2014

López-García, Margarita M. & Deloya, Cuauhtémoc 2022
2022
Loc

Tomarus rostratus

Dupuis, F. 2014: 1
2014
GBIF Dataset (for parent article) Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF