Cimbicidae, Curtis, 1825

Archibald, S. Bruce & Rasnitsyn, Alexandr P., 2023, Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands, Zootaxa 5278 (1), pp. 1-38 : 23-24

Cimbicidae subfamily incertae sedis

Genus and species incertae sedis

Fig. 22 .

Description. Forewing ( Fig. 22 ) length 15.5 mm, maximum width 5.4 mm, length/width 2.9; membrane lightly infuscate throughout, pterostigma dark, narrow, tapering, narrower than C-R before pterostigma; R+M shorter than 1-M; maximum length of cell 2r shorter than 3r (3r length about 1.6 2r); 2r-rs longer than Rs between 2r-m, 2r-rs, joining cell 3rm before its middle; 2r-m subvertical, short, slightly curved, joins Rs scarcely behind 2mcu; 1-M almost straight, sub-parallel to 1m-cu; 3r-m slightly angular toward base; cell 2mcu with M slightly shorter than 1m-cu; 2m-cu slightly curved toward base; cu-a postfurcal; anal cells meet at crossvein.

Remarks. The straight, short 2r-m excludes this specimen from the Cenocimbicinae . Its other forewing venation is a mixture of character states diagnostic of modern subfamilies: M+Sc-R junction is close to that of Rs+M as in Pachylostictinae and Corynidinae, but the anal veins are separate throughout, joined by a short crossvein 1a as in the Cimbicinae . We refrain from assigning this specimen to a subfamily or erecting a new one based on it, or naming it pending more specimens to clarify its position. We only note that this forewing is very different not only from those of all other Ypresian Cimbicidae , but from all others. Unfortunately, the wings of the only other known Ypresian non-cenocimbicine, the Green River Formation Eopachylosticta byrami (Cockerell) are too poorly preserved to compare.

Material. GSC 142844 , a forewing collected at McAbee by SBA ( GSC locality V-016800), May 19, 2001 ( Fig. 22 ), in the collections of the GSC (Vancouver).

Locality and age. McAbee, British Columbia, Canada, Hoodoo Face beds; mid-Ypresian.

Figures

FIGURE 22. GSC 142844, Cimbicidae subfamily, genus, and species incertae sedis: A, photograph; B, drawing.Scale bar 5 mm.