Daviesia incrassata Smith (1808b: 255)

Crisp, Michael D., Cayzer, Lindy, Chandler, Gregory T. & Cook, Lyn G., 2017, A monograph of Daviesia (Mirbelieae, Faboideae, Fabaceae), Phytotaxa 300 (1), pp. 448-450 : 448-450

108. Daviesia incrassata Smith (1808b: 255) , Bentham (1864: 83), Crisp (1984: 167), ( Crisp 1987a: 746), Crisp (1995: 1200), Wheeler et al. (2002: 746). Type: ‘Found at King George’s Sound, on the west coast of New Holland, by Mr. A. Menzies.’ Holotype: King George Sound, west coast of New Holland, (at 35, Menzies, 1803 (LINN); isotypes: BM (2 sheets), MO

Acacia dolabriformis H.L. Wendland (1820: 6) . Type: ‘ Nec flores nec fructus vidi, sed tantum unicum specimen in Nova Hollandia lectum ...’ Holotype: GOET.

Daviesia reversifolia Mueller (1859b: 145) , Bentham (1864: 83). Type: ‘In monte ‘Flat-top Hill’ juxta rivum Fitzgerald River. Mxw.’ Holotype: MEL 80382; isotypes: K,?MEL 80383.

Rounded to spreading shrubs, to 1 m high, glabrous, ± glaucous (to pruinose). Root anatomy with anomalous secondary thickening (cord type). Branchlets ascending or divaricate, gently to strongly flexuose, trigonous to terete, smooth when fresh, wrinkled-striate when dry. Phyllodes scattered, trigonous to terete, spreading at 45–90° or (in subsp. reversifolia ) retrorse, terete or vertically compressed or dilated, at least towards the base of the plant, inarticulate and sometimes decurrent, 5–15 mm long, smooth when fresh, wrinkled-striate when dry. Juvenile phyllodes vertically dilated and asymmetrically lobed (in subsp. incrassata ), or not retrorse, gently incurved, vertically compressed but not lobed (in subsp. reversifolia ), or strictly terete (in subsp. teres ), up to 8 cm long. Unit inflorescences 1(2) per axil, racemose, 2–4-flowered; peduncle occasionally somewhat triquetrous, 0.5–3 mm long; rachis flexuose, 0.25–2 mm long; subtending bracts spreading to ascending, oblong, to 1 mm long. Pedicels dilating towards the apex, terete, 2–4.5 mm long. Calyx 2.5–3 mm long including the 0.75–1 mm receptacle; lobes shallowly triangular, subequal with broad, shallow sinuses, upper 2 united higher than the lower 3, 0.5–1 mm long. Corolla : standard transversely elliptic to obovate to broadly so, emarginate, with a central groove, 7–8 × 6.5–8 mm including the 1.5–2 mm claw, orange with pink infusion towards the margin, deep red to blackish centrally and on radiating veins, fading to yellowish and grey; wings elliptic with a rounded apex, auriculate, 6–7.5 × 1.5–2.5 mm including the 1.5–2 mm claw, deep pink; keel half very broadly to transversely broadly ovate with an acute, beaked

A MONOGRAPH OF DAVIESIA

Phytotaxa 300 (1) © 2017 Magnolia Press • 241 apex, auriculate, 7–7.5 × 2 mm including the 3–4.5 mm claw, pink with a blackish tip. Stamens weakly dimorphic: inner whorl of 5 with longer, narrower filaments and shorter, subdorsifixed anthers; outer whorl of 5 with shorter filaments and longer, basifixed anthers; filaments all somewhat compressed, cohering towards base; anthers all 2- celled. Pod obliquely shallowly obtriangular with an acute to acuminate, beaked apex, turgid, 11–14 × 8–11 mm; upper suture undulate; lower suture acute. Seed not seen. ( Figs 109 , 110 ).

Flowering period:— June to November. Fruiting period: August to January.

Distribution:— Western Australia, widespread through the south-west from near Dongara to Mt Ragged, between the coast and the wetter (western and southern) fringes of the wheatbelt.

Affinity:— Daviesia incrassata is similar to D. brachyphylla , D. inflata , D. physodes and D. retrorsa . All four species are easily distinguished from D. incrassata by their articulate phyllode bases. Daviesia inflata also differs in having a bicoloured calyx, in which the margins of the lobes are distinctly paler than the rest. Daviesia retrorsa has phyllodes developed only towards the apex of the branchlets, smaller (9–11 mm long), less turgid pods, and the keel is neither incurved or beaked. Daviesia brevifolia has short, non-articulate phyllodes and is very similar to D. incrassata subsp. teres —see under the latter for distinguishing characters.

Figures

FIGURE 109. Daviesia incrassata subsp. incrassata. A. Flowering branchlet with terete upper phyllodes. B. Lower phyllode from the same plant, showing vertical dilation. C. Flowering branchlet with vertically compressed phyllodes. D. Lower phyllode from the same plant, showing pronounced vertical dilation. E. Inflorescence. F. Pod. A, B from Crisp 6132; C, D from Chapman (16)78; E from Crisp 1021; F from Crisp 5368. Drawn by A.L. Prowse. Adapted from Crisp (1995) with permission from CSIRO Publishing.

FIGURE 110. Daviesia incrassata subsp. reversifolia. A. Adult fruiting branchlet. B. Flowering branchlet showing developmental sequence of phyllodes from base (juveniles) to tip (adults). C. Juvenile phyllode enlarged. Intermediate between D. incrassata subsp. reversifolia and subsp. incrassata. D. Adult branchlet. E. Juvenile phyllode showing vertical dilation. Daviesia incrassata subsp. teres. F. Flowering adult branchlet. G. Juvenile branchlet. H. Pod. A from Crisp 4931; B, C from Bennett 919; D, E from Crisp 5095; F from Gittins 1686 (type); G from Crisp 5427; H from Chapman (108)77. Drawn by A.L. Prowse. Adapted from Crisp (1995) with permission from CSIRO Publishing.