Daviesia filipes Benth.

Crisp, Michael D., Cayzer, Lindy, Chandler, Gregory T. & Cook, Lyn G., 2017, A monograph of Daviesia (Mirbelieae, Faboideae, Fabaceae), Phytotaxa 300 (1), pp. 448-450 : 448-450

59. Daviesia filipes Benth. in Mitchell (1848: 363), Bentham (1864: 80), Stanley & Ross (1983: 254), Crisp (1995: 1193). Type: Not designated, but from the text one can determine: Collector—Mr. Drysdale, storekeeper. Date— between 30 May & 19 October 1846. Locality—near 2nd Depot, by R. Maranoa just W of Mt. Sowerby, ca. 26°13’S, 148°06’E. Lectotype (Crisp 1995: 1193): Subtropical New Holland, Camp 29, Lieut. Col. Sir T.L. Mitchell 558, 15 September 1846 (K, ex Herb. Bentham); isolectotype: BM, CGE (2 sheets), K (ex Herb. Hooker)

Daviesia longipes Domin (1926: 723) . Type: ‘ Queensland: Sandsteinhuegel der Dividing Range bei Pentland (DOMIN II. 1910).’ Lectotype (Crisp 1995: 1193): Queensland, in collibus arenaceis montium Dividing Range apud opp. Pentland, K. Domin 4442, February 1910 ( PR 527026 ). Syntype: same locality, K. Domin 4441, February 1910 ( PR 527025 ). Note: This is a trivial variant, differing from typical D. filipes only in being glabrous; the species varies considerably in the density of the indumentum and complete glabrousness is only one extreme.

Shrubs to 1.5 m high, hirsute, hispid or villous on vegetative parts, rarely glabrous. Root anatomy normal (unistelar). Branchlets terete, ascending or arching. Phyllodes fairly crowded, ascending at ca. 45°, mostly narrowly oblong, occasionally tapered to either end, rarely broader, apically acuminate and pungent, with recurved to ± flat margins, rounded to cuneate and articulate at the base, (6–)9–19 × 1.5–6 mm. Unit inflorescence 1 per axil, umbellate, 1(–3)-flowered, with or without compound, terminal, raceme- or umbel-like inflorescences, which are due to the suppression of the phyllodes, 1–5-flowered; peduncle 0.5–9 mm long; barren basal bracts forming an involucre, ca. 0.25 mm long; subtending bracts appressed, oblong, ca. 0.5 mm long. Pedicel 4–12 mm long, hooked. Calyx 2.5–3 mm long including the ca. 0.5 mm receptacle; upper 2 lobes united in a broad, truncate lip or united higher then the lower 3 and broadly triangular, ca. 1 mm long, scabrous; lower 3 lobes triangular, ca. 0.75 mm long. Corolla yellow with red markings or rarely pure yellow (Crisp 11731); standard broadly obovate to very broadly so, emarginate, 4.5–5 × 3.5–4 mm including the ca. 1.5 mm claw, yellow with 2 dull red basal blotches either side of the central, intensely yellow ‘eyes’; wings obovate with a rounded apex, auriculate, 4.5–5 × 1.5 mm including the 1–1.5 mm claw, yellow infused with dull maroon towards the base; keel half elliptic, acute, auriculate, saccate, 4.5–5 × 1.5 mm including the 1.5–2 mm claw, maroon. Stamens strongly dimorphic: inner whorl of 5 with longer, slender, terete filaments and shorter, round, versatile anthers with confluent thecae; outer whorl of 5 with shorter, broader, compressed filaments and longer, oblong, basifixed, 2-celled anthers; filaments free. Pod obliquely shallowly obtriangular, acuminate, somewhat compressed, 7–9 × 4–5 mm; upper suture sigmoid; lower suture ca. 90°. Seed globose to fusiform, 3.2–4 mm long, 1.7–2.2 mm broad, 1–1.4 mm thick, dark to light brown with or without black mottling; aril 1.2–1.7 mm long. ( Fig. 59 ).

138 • Phytotaxa 300 (1) © 2017 Magnolia Press

CRISP ET AL.

Flowering period:— April to November. Fruiting period: April to November.

Distribution:— Queensland, from drier inland slopes of the ranges of the wet tropics (e.g. Davies Creek and Windsor Tableland) south to Inglewood in the Darling Downs.

Habitat:— In eucalypt-dominated open forest or woodland on plateaux, ridges or moderate to steep slopes. Soils are sandy or loamy, often with gravel, and derived from sandstone or granite.

A MONOGRAPH OF DAVIESIA

Phytotaxa 300 (1) © 2017 Magnolia Press • 139

Affinity:— Daviesia pubigera is very similar to D. filipes , replacing it in New South Wales in the equivalent habitats of the western slopes of the Great Dividing Range. Daviesia pubigera differs in having an inflorescence rachis (the portion at base bearing bracts) that is hispid and proportionally longer, being about equal to or longer than the pedicel (portion above highest bract). Also, the flowers are somewhat larger in D. pubigera , e.g. the calyx including the stipe-like receptacle is 2.8–3.3 mm long. Daviesia umbellulata can have narrow phyllodes resembling those of D. filipes , and some plants have terminal inflorescences as in D. filipes subsp. terminalis . However D. umbellulata differs in having generally longer phyllodes (7–30 mm long), a greater number of flowers per unit inflorescence (3–6), shorter pedicels (4.5–5 mm long) and larger flowers (e.g. calyx 3–3.5 mm long, standard 5–7 × 5–7.5 mm).

Figures

FIGURE 59. Daviesia filipes subsp. filipes. A. Fertile branchlet. B. Flower. C. Pod. Daviesia filipes subsp. terminalis. D. Phyllode that is glabrous except along nerves. A–C from Crisp 2747; D from Staples 100874/4. Drawn by B-J. Osborne.