Trichoptera

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 6-7

publication ID

https://doi.org/ 10.5281/zenodo.210736

DOI

https://doi.org/10.5281/zenodo.5664460

persistent identifier

https://treatment.plazi.org/id/9F3E87DD-562B-FFD0-E89A-FF5AFA17F95D

treatment provided by

Plazi

scientific name

Trichoptera
status

 

Key to Fijian Trichoptera View in CoL families (adults) and Philopotamidae species (males)

1 Forewings shorter than 4.0 mm, narrow, pointed apically (figs. 6–10 in Marshall (1979)); antennae shorter than forewings; tho- rax with mesoscutellum pointed posteriorly (fig. 83 in Marshall (1979))................................................................................................. Hydroptilidae View in CoL (1 species: Oxyethira fijiensis Kelley, 1989 View in CoL ).

- Forewings longer than 4.0 mm, broad, rounded apically ( Fig. 5 View FIGURE 5 ); antennae as long as forewings or longer; thorax with mesoscutellum rounded or nearly straight posteriorly (figs. 53–54 in Marshall (1979))............................... 2

2(1’) Ocelli present........................................................................................ 3

- Ocelli absent......................................................................................... 4

3(2) Maxillary palps each with segment 2 swollen mesally and with apical segment flexible, distinctly longer than previous seg- ment............................................................. Philopotamidae View in CoL ( Chimarra View in CoL , 27 species)...8

- Maxillary palps each with segment 2 cylindrical and with apical segment not flexible, about as long as previous segment...................................................................... Hydrobiosidae View in CoL ( Apsilochorema View in CoL , 3 species).

4(2’) Maxillary palps each with apical segment flexible, distinctly longer than previous segment........................... 5

- Maxillary palps each with apical segment not flexible, about as long as previous segment........................... 6

5(4) Forelegs each with 2 spurs (preapical spur absent); thorax without mesonotal setal warts................................................................................................... Hydropsychidae View in CoL ( Abacaria View in CoL , 9 species).

- Forelegs each with 3 spurs (preapical spur present); thorax with pair of mesonotal setal warts......................................................................................... Polycentropodidae View in CoL ( Polyplectropus View in CoL , 8 species).

6(4’) Antennae each with scape distinctly longer than head; thorax with pair of distinct mesonotal setal warts, setae never in longitu- dinal rows..................................................................... Goeridae View in CoL ( Goera View in CoL , 3 species).

- Antennae each with scape shorter or about as long as head; thorax without mesonotal setal warts, setae always in pair of longi- tudinal rows.......................................................................................... 7

7(6’) Maxillary palps each 6-segmented; forewings less than 3x longer than broad... Calamoceratidae View in CoL ( Anisocentropus View in CoL , 5 species)

- Maxillary palps 5-segmented; forewings more than 3x longer than broad................................................................................ Leptoceridae View in CoL ( Oecetis View in CoL , 1 species; Triaenodes View in CoL , 3 species; Triplectides View in CoL , 1 species)

8(3) Hind wings each with centrally located, large, pale hyaline spot ( Figs. 10–13 View FIGURES 6 – 13 )..................................... 9

- Hind wings without large pale, hyaline spots ( Figs. 6–7 View FIGURES 6 – 13 ).................................................... 14

9(8) In genitalia, gonopods short, length about as long as high in lateral view ( Fig. 50 View FIGURES 50 – 54 ).............. C. schlingeri , new species.

- In genitalia, gonopods more elongate, longer than high in lateral view (cf. Figs. 60 View FIGURES 60 – 64 , 65 View FIGURES 65 – 69 , 70 View FIGURES 70 – 74 )......................... 10

10(9’) In genitalia, ventral branch of each gonopod more than 2x longer than wide in lateral view ( Fig. 60 View FIGURES 60 – 64 ) C. braueri , new species.

- In genitalia, ventral branch of each gonopod not as long as wide in lateral view ( Figs. 45 View FIGURES 45 – 49 , 65 View FIGURES 65 – 69 )........................ 11

11(10’) In genitalia, gonopods widest at mid-length in lateral view ( Fig. 70 View FIGURES 70 – 74 )..................... C. karoyanitensis , new species.

- In genitalia, gonopods widest beyond mid-length in lateral view ( Figs. 45 View FIGURES 45 – 49 , 55 View FIGURES 55 – 59 , 65 View FIGURES 65 – 69 )................................. 12

12(11’) In genitalia, dorsal branch of each gonopod forming ventrally pointing hook in lateral view ( Fig. 45 View FIGURES 45 – 49 ) C. signata Banks, 1936 View in CoL .

- In genitalia, dorsal branch of each gonopod forming posteriorly pointing triangle in lateral view ( Figs. 55 View FIGURES 55 – 59 , 65 View FIGURES 65 – 69 )......... 13

13(12’)In genitalia, dorsal margin of each gonopod nearly straight and as darkly sclerotized as posterior margin ( Fig. 55 View FIGURES 55 – 59 )........................................................................................... C. nathani , new species.

- In genitalia, dorsal margin of each gonopod strongly convex and more darkly sclerotized than posterior margin ( Fig. 65 View FIGURES 65 – 69 )...................................................................................... C. vitiensis , new species.

14(8’) Forewings each with large, pale hyaline spot............................................................... 15

- Forewings without large, pale hyaline spots............................................................... 16

15(14) Forewings each with large, pale hyaline spot located immediately anterior of posterior wing margin ( Fig. 6 View FIGURES 6 – 13 ); in genitalia, ter- gum X with prominent dorsal process on each lateral branch ( Fig. 35 View FIGURES 35 – 39 )....................... C. vanuensis , new species.

- Forewings each with large, pale hyaline spot located centrally in wings ( Fig. 7 View FIGURES 6 – 13 ); in genitalia, tergum X without dorsal pro- cesses ( Fig. 40 View FIGURES 40 – 44 )................................................................. C. macuatensis , new species.

16(14’)In genitalia, phallic apparatus without endothecal processes; phallotremal sclerite more than 2x longer than width of narrowest part of phallotheca ( Figs. 79 View FIGURES 75 – 79 , 84 View FIGURES 80 – 84 , 89 View FIGURES 85 – 89 )..................................................................... 17

- In genitalia, phallic apparatus with 1 or 2 pairs of endothecal processes; phallotremal sclerite less than 2x longer than width of narrowest part of phallotheca, or not discernable ( Figs. 139 View FIGURES 139 – 144 , 170 View FIGURES 170 – 174 )............................................... 19

17(16) In genitalia, gonopods acute apically in lateral view ( Fig. 75 View FIGURES 75 – 79 ); phallotremal sclerite more than 1/2 as long as phallotheca ( Fig. 79 View FIGURES 75 – 79 )............................................................................. C. tokotaai , new species.

- In genitalia, gonopods rounded apically in lateral view ( Figs. 80 View FIGURES 80 – 84 , 85 View FIGURES 85 – 89 ); phallotremal sclerite much less than 1/2 as long as phal- lotheca ( Figs. 84 View FIGURES 80 – 84 , 89 View FIGURES 85 – 89 ).................................................................................. 18

18(17’)In genitalia, segment IX rectangular and oblique, with ventral margin much shorter than height in lateral view ( Fig. 85 View FIGURES 85 – 89 ); each posterior margin produced into large, triangular plate immediately below its cercus ( Fig. 85 View FIGURES 85 – 89 ).. C. naitasirensis , new species.

- In genitalia, segment IX hyperbolic, with ventral margin subequal to height in lateral view ( Fig. 80 View FIGURES 80 – 84 ); each posterior margin without triangular plate below its cercus ( Fig. 80 View FIGURES 80 – 84 )............................................ C. vuda , new species.

19(16’)In genitalia, phallic apparatus with 2 pairs of endothecal processes ( Fig. 138 View FIGURES 134 – 138 )..................................... 20

- In genitalia, phallic apparatus with 1 pair of endothecal processes ( Fig. 169 View FIGURES 165 – 169 )..................................... 23

20(19) In genitalia, each lateral branch of tergum X with dorsal, rounded process ( Figs. 105 View FIGURES 105 – 110 , 111 View FIGURES 111 – 116 ).......................... 21

- In genitalia, each lateral branch of tergum X with dorsal, hook-shaped process ( Figs. 134 View FIGURES 134 – 138 , 139 View FIGURES 139 – 144 )...................... 22

21(20) In genitalia, anteroventral portion of segment IX rectangular in lateral view ( Fig. 105 View FIGURES 105 – 110 ); in tergum X, gap between dorsal and ventral branches nearly right-angled ( Fig. 105 View FIGURES 105 – 110 ); tergum X with triangular process on lateral margin in lateral view ( Fig. 105 View FIGURES 105 – 110 ); gonopods strongly expanded ventrally at base ( Fig. 105 View FIGURES 105 – 110 ); in phallus, endothecal processes no longer than as minimum breadth of phallotheca in ventral view ( Fig. 110 View FIGURES 105 – 110 )................................................. C. manni Banks, 1924 View in CoL .

- In genitalia, anteroventral portion of segment IX triangular in lateral view ( Fig. 111 View FIGURES 111 – 116 ); in tergum X, gap between dorsal and ventral branches wide-angled, nearly 180 degrees ( Fig. 111 View FIGURES 111 – 116 ); tergum X without triangular process on ventral margin in lateral view ( Fig. 111 View FIGURES 111 – 116 ); gonopods only slightly expanded ventrally at base ( Fig. 111 View FIGURES 111 – 116 ); in phallus, endothecal processes longer than minimum breadth of phallotheca in ventral view ( Fig. 116 View FIGURES 111 – 116 )............................... C. cakaudrovensis , new species.

22(20’) In genitalia, each dorsal branch of tergum X with darkly sclerotized apicodorsal hook ( Fig. 134 View FIGURES 134 – 138 ); each ventral branch of tergum X reaching as far posterior as each dorsal branch ( Fig. 134 View FIGURES 134 – 138 ); phallic apparatus without spicules ( Figs. 137, 138 View FIGURES 134 – 138 )............................................................................................ C. lavensis , new species.

- In genitalia, each dorsal branch of tergum X with weakly sclerotized subapicodorsal process ( Fig. 139 View FIGURES 139 – 144 ); each ventral branch of tergum X exceeding posteriorly each dorsal branch of tergum X ( Fig. 139 View FIGURES 139 – 144 ); phallic apparatus with group of spicules in addition to endothecal processes ( Figs. 143, 144 View FIGURES 139 – 144 )................................................ C. devoensis , new species.

23(19') In genitalia, tergum X with each lateral branch uniramus ( Figs. 90 View FIGURES 90 – 94 , 124 View FIGURES 124 – 128 , 155 View FIGURES 155 – 159 )..................................... 24

- In genitalia, tergum X with each lateral branch biramus ( Figs. 95 View FIGURES 95 – 99 , 117 View FIGURES 117 – 123 , 145 View FIGURES 145 – 149 )..................................... 29

24(23) In genitalia, gonopods clearly reaching beyond tergum X posteriorly ( Figs. 100 View FIGURES 100 – 104 , 155 View FIGURES 155 – 159 )........................... ... 25

- In genitalia, gonopods reaching as far out as tergum X or shorter ( Figs. 90 View FIGURES 90 – 94 , 150 View FIGURES 150 – 154 , 164 View FIGURES 160 – 164 ).............................. 26

25(24) In genitalia, segment IX long, with narrow anteroventral plates and straight posterior margin in lateral view ( Fig. 100 View FIGURES 100 – 104 ); each gonopod uniramus ( Fig. 100 View FIGURES 100 – 104 )....................................................... C. abacensis , new species.

- In genitalia, segment IX short, with broad anteroventral plates and convex posterior margin in lateral view ( Fig. 155 View FIGURES 155 – 159 ); each gonopod with slender dorsal branch and rounded ventral branch ( Fig. 155 View FIGURES 155 – 159 ).................. C. psychodida , new species.

26(24’) In genitalia, gonopods about as long as broad in lateral view ( Fig. 90 View FIGURES 90 – 94 )....................... C. veisarensis , new species.

- In genitalia, gonopods much shorter than broad in lateral view ( Fig. 124 View FIGURES 124 – 128 , 165 View FIGURES 165 – 169 , 170 View FIGURES 170 – 174 )................................ 27

27(26') In genitalia, distal part of each gonopod almost straight ( Figs. 124 View FIGURES 124 – 128 , 170 View FIGURES 170 – 174 )........................................ 33

- In genitalia, distal part of each gonopod curved posterad in lateral view ( Figs. 150 View FIGURES 150 – 154 , 165 View FIGURES 165 – 169 )............................ 28

28(27’) In genitalia, segment IX with slightly concave to straight posterior margin ( Fig. 150 View FIGURES 150 – 154 ); tergum X short, not reaching beyond gonopods in lateral view ( Fig. 150 View FIGURES 150 – 154 ); each lateral branch of tergum X with lateral subapical process ( Fig. 151 View FIGURES 150 – 154 ); gonopods each with rounded ventral branch in lateral view ( Fig. 150 View FIGURES 150 – 154 )..................................... C. tipulida , new species.

- In genitalia, segment IX with slightly convex posterior margin ( Fig. 165 View FIGURES 165 – 169 ); tergum X long, reaching well beyond gonopods in lateral view ( Fig. 165 View FIGURES 165 – 169 ); tergum X without lateral processes ( Fig. 166 View FIGURES 165 – 169 ); gonopods each with nearly rectangular ventral branch in lateral view ( Fig. 165 View FIGURES 165 – 169 ).............................................................. C. malickyi , new species.

29(23’) In genitalia, gonopods broad, nearly triangular in lateral view ( Fig. 117 View FIGURES 117 – 123 ).................... C. cartwrighti , new species.

- In genitalia, gonopods narrow, each with nearly parallel-sided dorsal branch ( Figs. 95 View FIGURES 95 – 99 , 129 View FIGURES 129 – 133 , 145 View FIGURES 145 – 149 , 160 View FIGURES 160 – 164 )................ 30

30(29’) In genitalia, tergum X with ventral branches produced posterad beyond dorsal branches ( Figs. 95 View FIGURES 95 – 99 , 145 View FIGURES 145 – 149 )................ 31

- In genitalia, tergum X with ventral branches not produced posterad beyond dorsal branches ( Figs. 129 View FIGURES 129 – 133 , 160 View FIGURES 160 – 164 )............ 32

31(30) In genitalia, apex of each dorsal and ventral branch of each gonopod parallel-sided and bent posterad C. fijiana , new species.

- In genitalia, apex of dorsal branch of each gonopod straight; dorsal branch nearly parallel-sided, ventral branch irregular ( Fig. 145 View FIGURES 145 – 149 ).......................................................................... C. helomyzida , new species.

32(30’) In genitalia, gonopods longer than segment IX in lateral view, slightly curved posterad along their length ( Fig. 129 View FIGURES 129 – 133 ); distal 1/2 of tergum X narrower than distal 3rd of gonopods in lateral view ( Fig. 129 View FIGURES 129 – 133 )................. C. kadavuensis , new species.

- In genitalia, gonopods shorter than segment IX in lateral view, strongly curved into 1/2-circle ( Fig. 160 View FIGURES 160 – 164 ); tergum X broader than distal 3rd of gonopods in lateral view ( Fig. 160 View FIGURES 160 – 164 )..................................... C. levuensis , new species.

33(27) In genitalia, posterolateral margins of segment IX each with posterior, setose lobe above its gonopod in lateral view ( Fig. 124 View FIGURES 124 – 128 ); gonopods each narrowing at 1/2-length into parallel-sided dorsal branch and with rounded ventral branch in lateral view ( Fig. 124 View FIGURES 124 – 128 ); tergum X with convex dorsal margin in lateral view ( Fig. 124 View FIGURES 124 – 128 ), divided into lateral processes in dorsal view ( Fig. 125 View FIGURES 124 – 128 ).................................................................................. C. kimminsi , new species.

- In genitalia, posterolateral margins of segment IX without posterior, setose lobes in lateral view ( Fig. 170 View FIGURES 170 – 174 ); gonopods each with dorsal branch narrowing distally along its full length and with ventral branch short, triangular in lateral view ( Fig. 170 View FIGURES 170 – 174 ); tergum X with stepped, concave dorsal margin in lateral view ( Fig. 170 View FIGURES 170 – 174 ), without lateral processes in dorsal view ( Fig. 171 View FIGURES 170 – 174 )................................................................................. C. obscurella Banks, 1924 View in CoL .

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Trichoptera

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