Phlegmariurus intermedius (Trevis.) B.Øllg., Rodriguésia

Øllgaard, Benjamin, Boudrie, Michel & Cremers, Georges, 2020, The Lycopodiaceae of Guyana, Suriname, and French Guiana, Phytotaxa 433 (2), pp. 101-134 : 117-118

6. Phlegmariurus intermedius (Trevis.) B.Øllg., Rodriguésia 63(2): 480. 2012. — Lycopodium intermedium Spring , in Martius, Fl. Bras. I , 2: 111. 1840, non Blume 1828.— Lycopodium reflexum Lam. var. intermedium (Trevis.) Baker, Handb. Fern-allies p. 11. 1887.— Urostachys intermedius (Spring) Herter, Repert. Sp. Nov. Regni Veg. 19: 163. 1923.— Urostachys commutatus Herter, Index Lyc. 57. 1949.— Huperzia intermedia Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874.—Synypes: BRAZIL. Rio de Janeiro: “Prope Rio de Janeiro ”, Freyreiss s.n. (BR [BR0000006965716]), Sellow s.n. (B). Fig. 9A .

Plants erect or ascending from a decumbent base, or spreading-scrambling over the ground with erect shoot apices, to 40 cm tall or to 80 cm long, at least to 10 times dichotomous, scrambling individuals often with strongly divergent dichotomies. Shoots homophyllous, equally thick throughout, (4–)5–7(–9) mm in diam. incl. leaves, sporangiate from ca. 10–30 cm above the base, often in separate periodically produced zones. Stems excl. leaves 1–1.5 mm thick at the base, tapering to ca 0.5–1 mm upward. Leaves borne in usually regular, alternating whorls of 5–7, these 0.5–1.5(–2) mm apart, forming 10–14 indistinct longitudinal ranks, usually uniformly strongly recurved and hook-like from an initially appressed leaf base, sometimes slightly recurved to perpendicular, linear-subulate to linear-lanceolate, widest at or just above the base, 4–6 × 0.5–0.7 mm, not twisted, adaxially slightly concave to slightly convex, with indistinct to slightly prominent vein, abaxially with strongly and broadly prominent vein, with smooth margins, firmly herbaceous to subcoriaceous, usually shining, with distinctly prominent decurrent leaf bases. Sporangia 1–1.3(–1.5) mm wide.

Distribution:—Guadeloupe, Dominican Republic?, Venezuela (BO-MO), Guyana, Brazil.

Habitats:—Terrestrial or rupestral on rock outcrops in the mountains.

Specimens examined:— GUYANA: s.loc., Schomburgk Ro. Add. Ser. s.n (HBG); Schomburgk Ri. 1192 (B, K, P [1192B]); Schomburgk Ro. ser. Roraima 31 (P).

Figures

FIGURE 9. A. Phlegmariurus intermedius—(Brazil, Harley et al. 22599,AAU).A. Growth habit.A1. Close-up of sporangiate division.— B. Phlegmariurus reflexus—(Brazil, Irwin et al. 29233, P). B. Growth habit. B1. Close-up of sporangiate division. Drawing by. K. Tind.