Euploca lagoensis (Warm.) Diane & Hilger, in Hilger and Diane, 2003

Feuillet, Christian, 2016, Two new combinations in Euploca Nutt. (Heliotropiaceae, Boraginales) and a conspectus of the species of the Guiana Shield area, PhytoKeys 61, pp. 101-124 : 106-108

Euploca lagoensis (Warm.) Diane & Hilger, in Hilger and Diane, 2003 Fig. 2

Euploca lagoensis (Warm.) Diane & Hilger, in Hilger and Diane: Bot. Jahrb. Syst. 125 (1): 48. 2003.

Schleidenia lagoensis Warm., Kjoeb. Vidensk. Meddel 1867: 15. 1868.

Euploca lagoensis Type: Brazil. Minas Gerais: Lagoa Santa ad ripam lacus, 5 Nov 1864, Warming s.n. (holotype C n.v. [scan!; photos F-21971, M, MO]; isotypes GH n.v. [fragment], P! [scan!]).

Schleidenia subracemosa Warm., Kjoeb. Vidensk. Meddel 1867: 15. 1868.

Euploca lagoensis Type: Brazil. Minas Gerais: Lagoa Santa, 28 Jan 1866, Warming s.n. (holotype C [photo F-21972, GH, US]; isotype P! [scan!]).

Heliotropium lagoense (Warm.) Gürke, in Engler and Prantl, Nat. Pflanzemf. 4(3a): 97. 1893. Type: Based on Schleidenia lagoensis Warm.

Heliotropium trinitense Urb., Symb. Ant. 7(3): 350. 1912.

Euploca lagoensis Type: Trinidad. in Piarco Savannah, 10 May 1895, Lunt 6030 (holotype B †; lectotype, designated by Förther 1998, pg. 225: S n.v. [scan!]; isotypes [fragments of holotype GH n.v. and lectotype GH n.v.]).

Type.

Based on Schleidenia lagoensis Warm.

Description.

Annual herbs, up to 15 cm tall, stems decumbent or prostrate, up to 30 cm long, glabrous or with scattered slender appressed trichomes. Leaves: petiole about 1 mm long, glabrous; lamina oblanceolate, 0.5-1.5 × 0.1-0.3 cm, apex acute to acuminate, base attenuate, margin entire, ciliate, adaxially glabrous, abaxially glabrous to puberulent. Flowers single, extra-axillary on leafy stems: pedicel up to 3 mm long, calyx of 5 lanceolate or cuneate, unequal lobes, 1.5-2.5 mm long at anthesis, puberulent in and out; corolla white, yellow at throat, tube swollen at base, 3-4 mm long, puberulent in and out, lobes ovate, 1-1.5 mm long, with sinus rounded, occasionally plaited with a minute lobule; stamens sessile, inserted about 0.5 mm from base, anthers joined, oblong, with an hairy apical appendage nearly as large as the anther; ovary obpyriform, about 0.5 mm diam.; style lacking or confused with the apex of the ovary; stigma sessile or subsessile, much larger at base; fruit rostrate, glabrous or nearly so, about 2 mm diam., obpyriform; nutlets ca. 1.5-2 mm long.

Distribution.

Venezuela ( Bolívar), Surinam, French Guiana, and Brazil (Amazonas); also Greater Antilles and from Mexico to Brazil and eastern Bolivia. Euploca lagoensis is new to French Guiana. I have seen only few specimens from the Guiana Shield.

Selected specimens studied.

Venezuela, Bolívar, near San Carlos, Laguna de la Culebra, 6 Apr 1925, fl, Pittier 11703 (P). Surinam, In 1885, W.F.R. Suringar s.n. (L); Mennega 907 (L). French Guiana, Savane Maillard, 6 km SE de Tonate-Macouria, 4°58'N, 52°26'W, 25 Nov 1999, Raynal-Roques and Jérémie 24694 (CAY, K, P, NY).

Note.

Melo and Semir (2010) cited the holotype as being at M. I do not think there is a specimen at M, only a photograph. Furthermore, if Warming worked in Munich, it was about 7 years after he described Schleidenia lagoensis . In the 1867 he was working in Copenhagen, I do not see any reason to reject Förther’s (1998) choice of the specimen at C as the holotype.