Scybalocanthon kelleri Pereira & Martínez, 1956

Silva, Fernando A. B. & Valois, Marcely, 2019, A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini), Zootaxa 4629 (3), pp. 301-341 : 323-324

Scybalocanthon kelleri Pereira & Martínez, 1956

( Figs. 1L , 4L , 5K , 6A )

Scybalocanthon kelleri Pereira & Martínez, 1956: 114 , 116 (original description and identification key); Vulcano & Pereira 1964: 638 (catalog); Vulcano & Pereira 1967: 554 (distribution, identification key); Halffter & Martínez 1977: 67 (species list).

Canthon kelleri: Krajcik 2006: 28 (catalog); Krajcik 2012: 63 (catalog).

Diagnosis. Specimens of S. kelleri ( Fig. 1L ) are similar to some specimens of S. sexspilotus ( Fig. 2C ) in having the pronotum bicolored, with a transverse black spot; aedeagus with parameres elongate, subrectangular, not excavated ventrally ( Figs. 4L, U ); and endophallus lacking bristles or microbristles close to the FLP sclerite ( Figs. 5K, U ). However, they can be distinguished by head, pygidium and femora uniformly colored.

Description. Body. Oval, lateral edges rounded. Surface completely microgranulate, with small and dense punctures randomly distributed. Color. Head and elytra opaque, pronotum with metallic sheen. Most of pronotum and elytra, lateral portion of hypomera, lateral portion of abdominal ventrites, and pygidium yellow or dark brown. Other parts black. Length. 7.7–9.0 mm. Thorax. Pronotum with transverse black spot in a butterfly or bat shape. Anterior angles of pronotum acute (approximately 80°). Lateral margin regularly curved outward, not forming an angle at the middle portion. Elytra. Striae thin and shiny, weakly impressed, punctures almost inconspicuous. Eighth stria with a thin carina at the anterior portion. Aedeagus. Parameres slightly asymmetrical, with subrectangular shape, truncate apically ( Fig. 4L ). Dorsal margin of parameres slightly curved inward from the basal to medial portions and slightly curved outward at the apical portion. Ventral margin of parameres substraight. SRP circular, with curved handle-shaped extension ( Fig. 5K ). FLP I-shaped ( Fig. 5K ). A+SA with two superposed and elongate sclerites ( Fig. 5K ). AS I-shaped ( Fig. 5K ).

Type material. Scybalocanthon kelleri Pereira & Martínez , holotype ♂ ( MZSP): “Sosomoco / Ost Columb.” // “ HOLOTYPUS ” / / “ Scybalocanthon / kelleri sp. n. ♂ / F. S. Pereira ” // “ Canthon / nigriceps / Har. / P. Pereira det. 951” // “10196” .

Non-type material [ 7 males, 5 females]: COLOMBIA: Sosomoco, [no date], [anonymous]— 1♂ ( CMNC); META, Quebrada Susamuco , 23 km W Villavicencio, 1000 m (4°12’4’’N, 73°47’40’’W), 3–5.iii.1972, S. & J. Peck, forest— 3♂ 4♀ ( CMNC); Vista Hermosa, Finca El Esfuerzo (73°35’42”W, 3°2’44”N), iv.1997, pitfall human feces, 200 m, A. Lopera— 1♂ ( CEMT); Buena Vista, i.1897, S. Burger— 1♂ ( CEMT) . ECUADOR: NAPO, Tena , 400–500 m (0°59’21’’S, 77°48’55’’W), 15–21.ii.1986, F. Génier, feces trap— 1♂ 1♀ ( CMNC) .

Distribution. Known from Colombia and Ecuador ( Fig. 6A ). Endemism areas: Brazilian sub-region: Pacific dominion: Sabana and Magdalena provinces; Boreal Brazilian dominion: Napo province (see Morrone 2014; fig. 12).

Remarks. “Sosomoco” is apparently an incorrect citation for quebrada Susumuco in the eastern portion of the Meta department in Colombia ( Maldonado-Ocampo et al. 2005; Salazar 2008).

Figures

FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm.

FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm.

FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm.

FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm.

FIGURE 6. Known distribution of species of Scybalocanthon.