Scorpiops harmsi, Kovařík, 2020

Kovařík, František, 2020, Nine new species of Scorpiops Peters, 1861 (Scorpiones: Scorpiopidae) from China, India, Nepal, and Pakistan, Euscorpius 302, pp. 1-43 : 10

Scorpiops harmsi sp. n.

( Figures 49–68 , 240 , Table 1) http://zoobank.org/urn:lsid:zoobank.org:act:1975A1CE-

FF6F-4811-9360-5AFE6C27AA92

TYPE LOCALITY AND TYPE REPOSITORY. Nepal, Annapurna massif, Mardi Himal (28.48°N 83.927°E), Deuvali, 2100 m a. s. l. ; FKCP.

TYPE MATERIAL. Nepal, Annapurna massif, Mardi Himal (28.48°N 83.927°E), Deuvali, 2100 m a. s. l., 10. V.2001, 1♀ (holotype), leg. J. Schmidt , FKCP.

ETYMOLOGY. The species epithet is a patronym honoring a German zoologist Danilo Harms, the curator of arachnids and myriapods at the Zoological Museum Hamburg, who greatly helped the author and his colleagues by loaning many type specimens of scorpions.

DIAGNOSIS (♀). Total length 55 mm long. Base color uniformly reddish brown to black, telson and tarsomere II of legs yellowish to reddish brown. Pectinal teeth number 5–6 in female, fulcra absent; pecten forms one compact unit with an incomplete furrow between places for marginal and middle lamellae. Patella of pedipalp with 17 (5 eb, 2 esb, 2 em, 4 est, 4 et) external and 7 ventral trichobothria. Chela of pedipalp with 4 ventral trichobothria located on ventral surface. Chelal trichobothrium Eb 3 located in proximal half of external surface between trichobothria Dt and Db. Fingers of pedipalps very slightly undulate in female. Chela length to width ratio 2.8 in female. Pedipalp movable finger with 40–42 IAD, which form a second row, parallel to MD (70–73 in number); there are also 4 ID and 7–8 OD present. Tarsomere II of legs with 7–8 stout median ventral spines in a row and two other parallel spines. Telson elongate and very sparsely granulate, length to depth ratio 3.31 in female.

DESCRIPTION (♀ holotype). Total length 55 mm. Male unknown. The habitus is shown in Figs. 49–50 . For position and distribution of trichobothria of pedipalps see Figs. 51–56 . Fingers of pedipalps are very slightly undulate in female ( Fig. 52 ).

Coloration ( Figs. 49–50 ). The base color is uniformly reddish brown to black, telson and tarsomere II of legs yellowish to reddish brown, sternites reddish black. Chelicerae are reddish brown and reticulate, fingers reddish brown.

Carapace and mesosoma ( Figs. 59–60 ). The entire carapace is covered with large and minute granules; carinae are absent. The anterior margin of the carapace is markedly depressed, convex in the middle. The carapace bears three lateral eyes. The mesosoma is granulated, with one median carina. Tergite VII is pentacarinate. The sternites are finely granulated with two parallel furrows except sternite VII, which bears four sparsely granulate carinae. Pectinal teeth number 7 in female, fulcra are absent. A pecten forms one compact unit with an incomplete furrow between places for marginal and middle lamellae.

Metasoma and telson ( Figs. 65–68 ). The metasoma is sparsely hirsute and granulated, with relatively large granules. Metasomal segment I with 10 carinae, II–IV with 8 carinae, and V with 5 carinae. The median lateral carinae of metasoma II are indicated by isolated granules that may coalesce into carinae. The dorsolateral carinae of segments III and IV granulate with sharp granules, which are not posteriorly terminated with a pronounced tooth. The telson is elongate and very sparsely granulate, with annular ring slightly indicated in female.

Pedipalps ( Figs. 51–58 ). The pedipalps are very sparsely hirsute. The patella bears 17 (5 eb, 2 esb, 2 em, 4 est, 4 et) external trichobothria and 7 ventral trichobothria. The chela bears 4 ventral trichobothria located on the ventral surface. The femur and patella are granulated. The femur has 5 granulose carinae and the patella has 5 complete carinae with reduced dorsal and ventral patellar spurs. The manus dorsally bears fine, rounded granules, which are not enlarged in the central area. The external surface of the chela is covered by minute and larger sparse granules. The movable fingers with 40–42 IAD, which form a second row, parallel to MD (70–73 in number); there are also 4 ID and 7–8 OD present.

Legs ( Figs. 61–64 ). The tibia and tarsomeres of legs with several setae not arranged into bristlecombs on dorsal surfaces but with rows of spines on dorsolateral and ventrolateral surface. Tarsomere II of legs I–IV with 7–8 stout median ventral spines in a row and two other parallel spines. The femur bears 3–4 and patella, 4–5 incomplete carinae; both femur and patella are finely granulated.

Measurements. See Table 1.

AFFINITIES. The described features distinguish S. harmsi sp. n. from all other species of the genus. The combination of five characters (chelal trichobothrium Eb 3 located in proximal half of external surface between trichobothria Dt and Db; fingers of pedipalps slightly undulate in female; pedipalp movable finger with 40–42 IAD; chela length to width ratio 2.8 in female; and patella of pedipalp with 7 ventral trichobothria) is unique in the entire genus Scorpiops .

DISTRIBUTION. Nepal ( Fig. 240 ).

Figures

Figure 240. Map of distribution of nine new species described here and Scorpiops vonwicki Birula, 1913.

Figures 49–50. Scorpiops harmsi sp. n., female holotype in dorsal (49) and ventral (50) views. Scale bar: 10 mm.

Figures 51–58. Scorpiops harmsi sp. n., female holotype, pedipalp segments. Chela dorsal (51), external (52) and ventral (53) views. Patella dorsal (54), external (55) and ventral (56) views. Movable finger dentition under white light (57) and UV fluorescence (58). Trichobothrial pattern is indicated by white circles.

Figures 59–68. Scorpiops harmsi sp. n., female holotype. Carapace and tergites I–III (59), posterior coxosternal area and sternites III–V (60). Left legs I–IV, retrolateral aspect (61–64 respectively). Telson lateral (65). Metasoma and telson lateral (66), ventral (67), and dorsal (68) views.