Luxiaria amasa ( Butler, 1878 )

Jiang, Nan, Xue, Dayong & Han, Hongxiang, 2014, A review of Luxiaria Walker and its allied genus Calletaera Warren (Lepidoptera, Geometridae, Ennominae) from China, Zootaxa 3856 (1), pp. 73-99 : 77-78

Luxiaria amasa ( Butler, 1878)

Figs 10–13 , 50 , 67 , 87 , 101

Bithia amasa Butler, 1878 , Ann. Mag. nat. Hist., (5) 1: 405. Holotype ♂, Japan: Yokohama (BMNH).

Luxiaria fasciosa Moore, 1888 , in Hewitson & Moore, Descr. new Indian lepid. Insects Colln late Mr W.S. Atkinson, 3: 254. Syntypes ♂, ♀, India: Darjeeling (BMNH).

Luxiaria fulvifascia Warren, 1894 , Novit. zool., 1: 440. Holotype ♂, Sumatra (BMNH).

Luxiaria contigaria amasa: Prout, 1915 , in Seitz, Macrolepid. World, 4: 350.

Luxiaria amasa: Wehrli, 1940 , in Seitz, Gross-Schmett. Erde, 4 (Suppl.): 407, pl. 33: b.

Diagnosis. The external characters of the species are similar to those of L. mitorrhaphes and L. acutaria , but L. amasa is distinguished by the following characters: the outer margin of the female forms a small protrusion at the end of M3; the dark bands outside the postmedial lines of both wings are reddish brown, while in L. mitorrhaphes and L. acutaria , those are greyish brown; reddish brown scales are present between the forewing submarginal line and the outer margin below M1 and the apex of hind wing, but are absent in L. mitorrhaphes and L. acutaria . The diagnostic characters of the genitalia can be seen under L. mitorrhaphes and L. acutaria .

Material examined. CHINA: Gausu ( IZCAS): 1♀, Kangxian, Qinghe Linchang, 1450–1650 m, 15.VII.1998, coll. Wang Shuyong. Zhejiang ( IZCAS): 3 ♂ 1♀, Taishun, Wuyanling, Shuangkengkou, 680 m, 29.VII.2005, coll. Lang Songyun; 1 ♂, Taishun, Shangfengxiang, 30.VII.2005, coll. Lang Songyun. Hubei ( IZCAS): 8 ♂, Xingshan, Longmenhe, 1300–1350 m, 21.VI–14.VII.1993, 6 –9.V.1994, 8 –14.IX.1994, coll. Song Shimei & Li Wenzhu; 3♀, Shennongjia, Dajiuhu, 1800 m, 1.VIII.1981, coll. Han Yinheng; 2♀, Shennongjia, Songluo, 920 m, 31.VIII.1981, coll. Han Yinheng. Jiangxi ( IZCAS): 4♀ Doushui, 29.VI–5.VII.1975, coll. Song Shimei; 2 ♂ 4♀, Jiulianshan, 9–14.VI.1975, coll. Zhang Baolin & Song Shimei. Hunan ( IZCAS): 1 ♂, Zhangjiajie, 8.X.1988; 1 ♂ 1♀, Zhangjiajie, Suoxiyu, 400 m, 17.X.1988. Fujian ( IZCAS): 20 ♂ 4♀, Wuyishan, Sangang, 700–704 m, 11.VIII.1979, 22–25.IV.1982, 30.VI.1982, 13–18.IX.1982, 10.V–6.VI.1983, 13.VIII.1983, 20.X.2005, 3.VI.2006, 11 –14.VIII.2009, coll. Xue Dayong et al.; 2♀, Wuyishan, Dazhulan, 1150 m, 28.VII.2006, coll. Yang Chao; 2 ♂, Jianyang, Huangkeng, Aotou, 750–950 m, 30.IV–1.V.1960, coll. Zhang Yiran & Zuo Yong; 1♀, Guadun, 17.VI.1981, coll. Jiang Fan; 1 ♂, Wuping, Liangyeshan, Kongxia, 480–627 m, 17–19.XI.2008, coll. Chen Fuqiang; 1♀, Nanping, Dafeng, 11.VI.1981. Hainan ( IZCAS): 1♀, Baisha, Nankai, Nanmaola, 1261 m, 10.V.2009, coll. Chen Fuqiang; 1♀, Qiongzhong, Limuling, 620 m, 15.V.2007, coll. Lang Songyun; 3♀, Jianfengling, 12.IV.1982, 18.V.1982, 8.VII.1982, coll. Liu Yuanfu et al. Guangxi ( IZCAS): 2 ♂ 1♀, Napo, Defu, 1350 m, 9–18.VI.2000, coll. Yao Jian & Li Wenzhu; 1 ♂ 1♀, Jinxiu, Luoxiang, 200–400 m, 15.V.1999, coll. Han Hongxiang & Li Wenzhu; 2 ♂ 1♀, Longsheng, 10–14.VI.1980, coll. Wang Linyao; 1 ♂ 1♀, Miao’ershan, Jiuniuchang, 1150–1600 m, 7–15.VII.1985, coll. Fang Chenglai; 1 ♂, Fangcheng, Fulong, 500 m, 24.V.1999, coll. Zhang Yanzhou; 1♀, Jinxiu, Linhai Shanzhuang, 1000 m, 2.VII.2000, coll. Li Wenzhu. Sichuan ( IZCAS): 1 ♂ 2♀, Emeishan, Qingyinge, 800–1000 m, 18–20.IV.1957, 4.VII.1957, coll. Huang Keren &Wang Zongyuan; 1♀, Emeishan, 2100 m, 25.VI.1955, coll. Yang Xingchi; 4 ♂, Wanxian, Wang’erbao, 1200 m, 10.VII.1993, 12.VIII.1993, 28.V.1994, coll. Song Shimei & Li Wenzhu; 3 ♂, Fengdu, Shiping, 610 m, 2.VI.1994, coll. Li Wenzhu; 1♀, Xichuang, 4.VIII.1974, coll. Han Yinheng; 1 ♂, Changshou, Nanmuyuan, 450 m, 9.VI.1994, coll. Li Wenzhu; 1 ♂, Ya’an, Sichuan Agricultural University, 580 m, 13.V.2009, coll. Li Jing; 2 ♂, Qingchengshan, 1000 m, 22.V–4.VI.1979, coll. Gao Ping. Chongqing ( IZCAS): 1 ♂, Jinyunshan, 800 m, 14.VI.1994, coll. Li Wenzhu. Yunnan ( IZCAS): 9 ♂ 3♀, Tengchong, Dahaoping, 2020 m, 5–7.VIII.2007, coll. Xue Dayong et al.; 2 ♂, Tengchong, Heinitang, 1930 m, 28–30.V.1992, coll. Xue Dayong; 8 ♂ 2♀, Baoshan, Baihualing, 1520 m, 11–13.VIII.2007, coll. Wu Chunguang & Lang Songyun; 1 ♂, Yongsheng, Liude, 2250 m, 10.VII.1984, coll. Liu Dajun; 1 ♂, Lijiang, Yuhu, 2700 m, 27.VII.1984, coll. Liu Dajun; 1 ♂, Ruili, Dengga, 980 m, 6–8.VI.1992, coll. Xue Dayong; 2 ♂, Yinghong, Dongfeng Famuchang, 24.IV.1978. Tibet ( IZCAS): 3 ♂ 9♀, Mêdog, 1091 m, 20–23.VIII.2006, coll. Lang Songyun; 1 ♂ 1♀, Mêdog, Road from Bomi to Mêdog, 80 Km, 2118 m, 24–25.VIII.2006, coll. Lang Songyun; 1 ♂, Mêdog, Aniqiao, 1060 m, 12–13.VIII.2006, coll. Lang Songyun; 8 ♂ 1♀, Bomi, Yi’ong, Tangmai, 2079–2300 m, 10.VIII.1983, 29–31.VIII.2005, 29–30.VIII.2006, coll. Han Yinheng et al.; 8 ♂ 2♀, Nyingchi, Pêlung, 2115 m, 1–2.IX.2005, coll. Wang Xuejian; 15 ♂, Zayü, Shang Zayü, 1534–1960 m, 21–26.VIII.2005, coll. Wang Xuejian; 1 ♂, Zham, 2200 m, 25.VI.1975, coll. Wang Ziqing.

Distribution. China (Gansu, Zhejiang, Hubei, Jiangxi, Hunan, Fujian, Taiwan, Guangdong, Hainan, Hong Kong, Guangxi, Sichuan, Chongqing, Yunnan, Tibet), Russia, Japan, Korean Peninsula, India, Nepal, Indonesia (Borneo, Java).

Remarks. Populations of L. amasa from China belong to the nominate subspecies.

Figures

FIGURES 1 – 28. Adults. 1 – 5. Luxiaria mitorrhaphes. 1, male (with a black patch at the anal margin of forewing, Zhejiang); 2, ditto, underside; 3, male (without a black patch at the anal margin of forewing, Zhejiang); 4, female (Fujian); 5, ditto, underside; 6 – 9. L. acutaria. 6, male (Tibet); 7, ditto, underside; 8, female (Tibet); 9, ditto, underside; 10 – 13. L. amasa. 10, male (Tibet); 11, ditto, underside; 12, female (Yunnan); 13, ditto, underside; 14 – 20. L. emphatica. 14, male (holotype, India, BMNH); 15, male (syntype of Luxiaria versiformis, Peninsular Malaysia, BMNH); 16, male (holotype of Luxiaria costinota, Taiwan, BMNH); 17, male (Guangxi); 18, ditto, underside; 19, female (Hainan); 20, ditto, underside; 21 – 24. L. tephrosaria. 21, male (Yunnan); 22, ditto, underside; 23, female (Yunnan); 24, ditto, underside; 25 – 28. L. phyllosaria. 25, male (Guangxi); 26, ditto, underside; 27, female (Hainan); 28, ditto, underside. Scale bar = 1 cm.

FIGURES 48 – 53. Male genitalia. 48, Luxiaria mitorrhaphes; 49, L. acutaria; 50, L. amasa; 51, L. emphatica (holotype, BMNH); 52, Luxiaria emphatica (holotype of Luxiaria costinota, BMNH); 53, L. emphatica. Scale bar = 1 mm.

FIGURES 65 – 76. Aedeagus. 65, Luxiaria mitorrhaphes; 66, L. acutaria; 67, L. amasa; 68, L. emphatica (holotype, BMNH); 69, Luxiaria emphatica (holotype of Luxiaria costinota, BMNH); 70, L. emphatica; 71, L. tephrosaria; 72, L. phyllosaria; 73, Calletaera obliquata; 74, C. obvia sp. nov. (paratype); 75, C. acuticornuta sp. nov. (paratype); 76, C. rotundicornuta sp. nov. (paratype). Scale bar = 1 mm.

FIGURES 77 – 98. Aedeagus and sternites of of male and female abdomen. 77 – 81. Aedeagus. 77, Calletaera consimilaria; 78, C. subexpressa; 79, C. postvittata (BMNH); 80, C. dentata sp. nov. (holotype); 81, C. trigonoprocessus sp. nov. (holotype). Scale bar = 1 mm. 82 – 84. Eighth sternite of male abdomen. 82, Luxiaria phyllosaria; 83, Calletaera obliquata; 84, C. subexpressa. 85 – 98. Seventh sternite of female abdomen. 85, Luxiaria mitorrhaphes; 86, L. acutaria; 87, L. amasa; 88, L. emphatica; 89, L. tephrosaria; 90, L. phyllosaria; 91, Calletaera obliquata; 92, C. obvia sp. nov. (paratype); 93, C. acuticornuta sp. nov. (paratype); 94, C. rotundicornuta sp. nov. (paratype); 95, C. consimilaria; 96, C. subexpressa; 97, C. dentata sp. nov. (paratype); 98, C. trigonoprocessus sp. nov. (paratype).

FIGURES 99 – 106. Female genitalia. 99, Luxiaria mitorrhaphes; 100, L. acutaria; 101, L. amasa; 102, L. emphatica; 103, L. tephrosaria; 104, L. phyllosaria; 105, Calletaera obliquata; 106, C. obvia sp. nov. (paratype). Scale bar = 1 mm.